andthereareindicationsthattheplanetsdifferfromoneanotherintheir
  properheats,aswellasintheheatswhichtheyreceivefromthesun。Bearing
  inmindthattheyalsodifferintheinclinationsoftheirorbits,theinclinations
  oftheiraxes,intheirspecificgravities,andintheirphysicalconstitutions,
  weseehowdecidedisthecomplexitywroughtintheSolarSystembythosesecondaryredistributionswhichhaveaccompaniedtheprimaryredistribution。§118。Passingfromillustrations,which,asassumingthenebular
  hypothesis,mustbeclassedasmoreorlesshypothetical,letusdescendtoevidencelessopentoobjection。ItisnowagreedamonggeologiststhattheEarthwasonceamoltenmass。
  Originally,then,itwascomparativelyhomogeneousinconsistence;and,because
  ofthecirculationwhichtakesplaceinheatedliquids,musthavebeencomparatively
  homogeneousintemperature。Itmust,too,havebeensurroundedbyanatmosphere
  consistingpartlyoftheelementsofairandwater,andpartlyofthosevarious
  otherelementswhichassumegaseousformsathightemperatures。Coolingby
  radiationmust,afteranimmensetime,haveresultedindifferentiatingthe
  portionmostabletopartwithitsheat;namely,thesurface。Afurthercooling,
  leadingtodepositionofallsolidifiableelementscontainedintheatmosphere,
  andthentoprecipitationofthewater,leavingbehindtheair,mustthus
  havecausedasecondmarkeddifferentiation;andasthecondensationcommenced
  onthecoolestpartsofthesurface—namely,aboutthepolestheremustsohaveresultedthefirstgeographicaldistinctions。Totheseillustrationsofgrowingheterogeneity,inferredfromknownlaws,
  Geologyaddsanextensiveseriesthathavebeeninductivelyestablished。
  TheEarth'sstructurehasbeenageafteragefurthercomplicatedbyadditions
  tothestratawhichformitscrust;andithasbeenageafteragemademore
  variousbytheincreasingcompositionofthesestrata;themorerecentof
  which,formedfromthedetritusofthemoreancient,aremanyofthemrendered
  highlycomplexbythemixturesofmaterialstheycontain。Thisheterogeneity
  hasbeenvastlyaugmentedbytheactionsoftheEarth'snucleusonitsenvelope;
  whencehaveresultednotonlymanykindsofigneousrocks,butthetilting
  upofsedimentarystrataatallangles,theformationoffaultsandmetallic
  veins,theproductionofendlessdislocationsandirregularities。Again,
  geologiststeachusthattheEarth'ssurfacehasbeengrowingmorevaried
  inelevation——thatthemostancientmountain—systemsarethesmallest,
  andtheAndesandHimalayasthemostmodern;while,inallprobability,there
  havebeencorrespondingchangesinthebedoftheocean。Asaconsequence
  ofthisceaselessmultiplicationofdifferences,wenowfindthatnoconsiderable
  portionoftheEarth'sexposedsurface,islikeanyotherportion,eitherincontour,ingeologicstructure,orinchemicalcomposition。Therehasbeensimultaneouslygoingonagradualdifferentiationofclimates。
  AsfastastheEarthcooledanditscrustsolidified,inequalitiesoftemperature
  arosebetweenthosepartsofitssurfacemostexposedtotheSunandthose
  lessexposed;andthusintimetherecametobethemarkedcontrastsbetween
  regionsofperpetualiceandsnowregionswherewinterandsummeralternately
  reignforperiodsvaryingaccordingtothelatitude,andregionswheresummer
  followssummerwithscarcelyanappreciablevariation。Meanwhile,elevations
  andsubsidences,recurringhereandthereovertheEarth'scrust,andproducing
  irregulardistributionsoflandandsea,haveentailedvariousmodifications
  ofclimatebeyondthosedependentonlatitude;whileayetfurtherseries
  ofsuchmodificationshasbeencausedbyincreaseddifferencesofheight
  inthesurface,whichinsundryplaceshavebroughtarctic,temperate,and
  tropicalclimatestowithinafewmilesofoneanother。Thegeneralresults
  are,thateveryextensiveregionhasitsownmeteorologicalconditions,and
  thateverylocalityineachregiondiffersmoreorlessfromothersinthoseconditions:asalsoinitsstructure,itscontour,itssoil。ThusbetweenourexistingEarth,thephenomenaofwhosevariedcrustneither
  geographers,geologists,mineralogists,normeteorologistshaveyetenumerated,
  andthemoltenglobeoutofwhichitwasevolved,thecontrastinheterogeneityisstriking。§119。Theclearest,mostnumerous,andmostvariedillustrations
  oftheadvanceinmultiformitythataccompaniestheadvanceinintegration,
  arefurnishedbylivingbodies。Distinguishedasthesearebythegreatquantity
  oftheircontainedmolecularmotion,theyexhibitinanextremedegreethe
  secondaryre—distributionswhichcontainedmotionfacilitates。Thehistory
  ofeveryplantandeveryanimal,whileitisahistoryofincreasingbulk,
  isalsoahistoryofsimultaneously—increasingdifferencesamongtheparts。Thistransformationhasseveralaspects。Thechemicalcompositionwhichisalmostuniformthroughoutthesubstance
  ofagerm,vegetaloranimal,graduallyceasestobeuniform。Theseveral
  compounds,nitrogenousandnon—nitrogenous,whichwerehomogeneouslymixed,
  segregatebydegrees,becomediverselyproportionedindiverseplaces,and
  producenewcompoundsbytransformationormodification。Inplantsthealbuminous
  andamylaceousmatterswhichformthesubstanceoftheembryo,giveorigin
  heretoapreponderanceofchlorophyllandtheretoapreponderanceofcellulose。
  Overthepartsthatarebecomingleaf—surfaces,certainofthematerials
  aremetamorphosedintowax。Inthisplacestarchpassesintooneofitsisomeric
  equivalents,sugar;andinthatplaceintoanotherofitsisomericequivalents,
  gum。Bysecondarychangesomeofthecelluloseismodifiedintowood;while
  someofitismodifiedintothealliedsubstancewhich,inlargemasses,
  wecallcork。Andthemorenumerouscompoundsthusarising,initiatefurther
  unlikenessesbyminglinginunlikeratios。Theyelk,oressentialpartof
  ananimal—ovum,havingcomponentswhichareatfirstevenlydiffusedamong
  oneanother,chemicallytransformsitselfinlikemanner。Itsprotein,its
  fats,itssalts,becomedissimilarlyproportionedindifferentlocalities;
  andmultiplicationofisomericformsleadstofurthermixturesandcombinations
  thatconstituteminordistinctionsofparts。Hereamassdarkeningbyaccumulation
  ofhematine,presentlydissolvesintoblood。Therefattyandalbuminousmatters
  uniting,composenerve—tissue。Atthisspotthenitrogenoussubstancetakes
  onthecharacterofcartilage;andatthat,calcareoussalts,gatheringtogether
  inthecartilage,laythefoundationofbone。Allthesechemicaldifferentiationsslowlybecomemoremarkedandmorenumerous。Simultaneouslyarisecontrastsofminutestructure。Distincttissuestake
  theplaceofmatterthathadpreviouslynorecognizableunlikenessesofparts;
  andeachofthetissuesfirstproducedundergoessecondarymodifications,
  causingsub—speciesoftissues。Thegranularprotoplasmofthevegetalgerm,
  equallywiththatwhichformstheunfoldingpointofeveryshoot,givesorigin
  tocellsthatareatfirstalike。Someofthese,astheygrow,flattenand
  unitebytheiredgestoformtheouterlayer。Otherslengthen,andatthe
  sametimejointogetherinbundlestolaythefoundationofwoody—fibre。
  Beforemuchelongating,certainofthesecellsshowabreaking—upofthe
  liningdeposit,which,duringelongation,becomesaspiralthread,orareticulated
  framework,oraseriesofrings;andbythelongitudinalunionofcellsso
  lined,vesselsareformed。Meanwhileeachofthesedifferentiatedtissues
  isre—differentiated:instancethatconstitutingtheessentialpartofa
  leaf,theupperstratumofwhichiscomposedofchlorophyll—cellsremaining
  closelypacked,whilethelowerstratumbecomesspongy。Ofthesamegeneral
  characterarethetransformationsundergonebythefertilizedovum,which,
  atfirstaclusterofsimilarcells,quicklyreachesastagemarkedbydissimilarity
  ofthecells。Morefrequentlyrecurringfissionofthesuperficialcells,
  aresultingsmallersizeofthem,andsubsequentunionofthemintoanouter
  layer,constitutethefirstdifferentiation;andthemiddleareaofthis
  layerisrenderedunliketherestbystillmoreactiveprocessesoflike
  kind。Bysuchmodificationsuponmodifications,manyandvarious,arisethe
  classesandsub—classesoftissueswhich,intricatelycombinedonewithanother,composeorgans。Equallyconformingtothelawarethechangesingeneralshapeandin
  theshapesoforgans。Allgermsareatfirstspheresandalllimbsareat
  firstbudsormereroundedlumps。Fromthisprimordialuniformityandsimplicity,
  theretakeplacedivergences,bothofthewholesandoftheleadingparts,
  towardsmultiformityofcontourandtowardscomplexityofcontour。Remove
  thecompactly—foldedyoungleavesthatterminateeveryshoot,andthenucleus
  isfoundtobeacentralknobbearinglateralknobs,oneofwhichmaygrow
  intoeitheraleaf,asepal,apetal,astamen,oracarpel:alltheseeventually
  ——unlikepartsbeingatfirstalike。Theshootsthemselvesalsodepartfrom
  theirprimitiveunityofform;andwhileeachbranchbecomesmoreorless
  differentfromtherest,thewholeexposedpartoftheplantbecomesdifferent
  fromtheimbeddedpart。So,too,isitwiththeorgansofanimals。Oneof
  theArthropoda,forinstance,haslimbsthatwereoriginallyindistinguishable
  fromoneanother——composedahomogeneousseries;butbycontinuousdivergences
  therehavearisenamongthemunlikenessesofsizeandform,suchaswesee
  inthecrabandthelobster。Vertebratecreaturesequallyexemplifythis
  truth。Thewingsandlegsofabirdareofsimilarshapeswhentheybud—outfromthesidesoftheembryo。Thusineveryplantandanimal,conspicuoussecondaryre—distributions
  accompanytheprimaryre—distribution。Afirstdifferencebetweentwoparts;
  ineachofthesepartsotherdifferenceswhichpresentlybecomeasmarked
  asthefirst;andalikemultiplicationofdifferencesingeometricalprogression,
  untilthereisreachedthatcomplexcombinationconstitutingtheadult。This
  isthehistoryofalllivingthingswhatsoever。PursuinganideawhichHarvey
  setafloat,ithasbeenshownbyWolffandVonBaer,thatduringitsdevelopment
  eachorganismpassesfromastateofhomogeneitytoastateofheterogeneity。Foragenerationthistruthhasbeenacceptedbybiologists。*