andthereareindicationsthattheplanetsdifferfromoneanotherintheir
properheats,aswellasintheheatswhichtheyreceivefromthesun。Bearing
inmindthattheyalsodifferintheinclinationsoftheirorbits,theinclinations
oftheiraxes,intheirspecificgravities,andintheirphysicalconstitutions,
weseehowdecidedisthecomplexitywroughtintheSolarSystembythosesecondaryredistributionswhichhaveaccompaniedtheprimaryredistribution。§118。Passingfromillustrations,which,asassumingthenebular
hypothesis,mustbeclassedasmoreorlesshypothetical,letusdescendtoevidencelessopentoobjection。ItisnowagreedamonggeologiststhattheEarthwasonceamoltenmass。
Originally,then,itwascomparativelyhomogeneousinconsistence;and,because
ofthecirculationwhichtakesplaceinheatedliquids,musthavebeencomparatively
homogeneousintemperature。Itmust,too,havebeensurroundedbyanatmosphere
consistingpartlyoftheelementsofairandwater,andpartlyofthosevarious
otherelementswhichassumegaseousformsathightemperatures。Coolingby
radiationmust,afteranimmensetime,haveresultedindifferentiatingthe
portionmostabletopartwithitsheat;namely,thesurface。Afurthercooling,
leadingtodepositionofallsolidifiableelementscontainedintheatmosphere,
andthentoprecipitationofthewater,leavingbehindtheair,mustthus
havecausedasecondmarkeddifferentiation;andasthecondensationcommenced
onthecoolestpartsofthesurface—namely,aboutthepolestheremustsohaveresultedthefirstgeographicaldistinctions。Totheseillustrationsofgrowingheterogeneity,inferredfromknownlaws,
Geologyaddsanextensiveseriesthathavebeeninductivelyestablished。
TheEarth'sstructurehasbeenageafteragefurthercomplicatedbyadditions
tothestratawhichformitscrust;andithasbeenageafteragemademore
variousbytheincreasingcompositionofthesestrata;themorerecentof
which,formedfromthedetritusofthemoreancient,aremanyofthemrendered
highlycomplexbythemixturesofmaterialstheycontain。Thisheterogeneity
hasbeenvastlyaugmentedbytheactionsoftheEarth'snucleusonitsenvelope;
whencehaveresultednotonlymanykindsofigneousrocks,butthetilting
upofsedimentarystrataatallangles,theformationoffaultsandmetallic
veins,theproductionofendlessdislocationsandirregularities。Again,
geologiststeachusthattheEarth'ssurfacehasbeengrowingmorevaried
inelevation——thatthemostancientmountain—systemsarethesmallest,
andtheAndesandHimalayasthemostmodern;while,inallprobability,there
havebeencorrespondingchangesinthebedoftheocean。Asaconsequence
ofthisceaselessmultiplicationofdifferences,wenowfindthatnoconsiderable
portionoftheEarth'sexposedsurface,islikeanyotherportion,eitherincontour,ingeologicstructure,orinchemicalcomposition。Therehasbeensimultaneouslygoingonagradualdifferentiationofclimates。
AsfastastheEarthcooledanditscrustsolidified,inequalitiesoftemperature
arosebetweenthosepartsofitssurfacemostexposedtotheSunandthose
lessexposed;andthusintimetherecametobethemarkedcontrastsbetween
regionsofperpetualiceandsnowregionswherewinterandsummeralternately
reignforperiodsvaryingaccordingtothelatitude,andregionswheresummer
followssummerwithscarcelyanappreciablevariation。Meanwhile,elevations
andsubsidences,recurringhereandthereovertheEarth'scrust,andproducing
irregulardistributionsoflandandsea,haveentailedvariousmodifications
ofclimatebeyondthosedependentonlatitude;whileayetfurtherseries
ofsuchmodificationshasbeencausedbyincreaseddifferencesofheight
inthesurface,whichinsundryplaceshavebroughtarctic,temperate,and
tropicalclimatestowithinafewmilesofoneanother。Thegeneralresults
are,thateveryextensiveregionhasitsownmeteorologicalconditions,and
thateverylocalityineachregiondiffersmoreorlessfromothersinthoseconditions:asalsoinitsstructure,itscontour,itssoil。ThusbetweenourexistingEarth,thephenomenaofwhosevariedcrustneither
geographers,geologists,mineralogists,normeteorologistshaveyetenumerated,
andthemoltenglobeoutofwhichitwasevolved,thecontrastinheterogeneityisstriking。§119。Theclearest,mostnumerous,andmostvariedillustrations
oftheadvanceinmultiformitythataccompaniestheadvanceinintegration,
arefurnishedbylivingbodies。Distinguishedasthesearebythegreatquantity
oftheircontainedmolecularmotion,theyexhibitinanextremedegreethe
secondaryre—distributionswhichcontainedmotionfacilitates。Thehistory
ofeveryplantandeveryanimal,whileitisahistoryofincreasingbulk,
isalsoahistoryofsimultaneously—increasingdifferencesamongtheparts。Thistransformationhasseveralaspects。Thechemicalcompositionwhichisalmostuniformthroughoutthesubstance
ofagerm,vegetaloranimal,graduallyceasestobeuniform。Theseveral
compounds,nitrogenousandnon—nitrogenous,whichwerehomogeneouslymixed,
segregatebydegrees,becomediverselyproportionedindiverseplaces,and
producenewcompoundsbytransformationormodification。Inplantsthealbuminous
andamylaceousmatterswhichformthesubstanceoftheembryo,giveorigin
heretoapreponderanceofchlorophyllandtheretoapreponderanceofcellulose。
Overthepartsthatarebecomingleaf—surfaces,certainofthematerials
aremetamorphosedintowax。Inthisplacestarchpassesintooneofitsisomeric
equivalents,sugar;andinthatplaceintoanotherofitsisomericequivalents,
gum。Bysecondarychangesomeofthecelluloseismodifiedintowood;while
someofitismodifiedintothealliedsubstancewhich,inlargemasses,
wecallcork。Andthemorenumerouscompoundsthusarising,initiatefurther
unlikenessesbyminglinginunlikeratios。Theyelk,oressentialpartof
ananimal—ovum,havingcomponentswhichareatfirstevenlydiffusedamong
oneanother,chemicallytransformsitselfinlikemanner。Itsprotein,its
fats,itssalts,becomedissimilarlyproportionedindifferentlocalities;
andmultiplicationofisomericformsleadstofurthermixturesandcombinations
thatconstituteminordistinctionsofparts。Hereamassdarkeningbyaccumulation
ofhematine,presentlydissolvesintoblood。Therefattyandalbuminousmatters
uniting,composenerve—tissue。Atthisspotthenitrogenoussubstancetakes
onthecharacterofcartilage;andatthat,calcareoussalts,gatheringtogether
inthecartilage,laythefoundationofbone。Allthesechemicaldifferentiationsslowlybecomemoremarkedandmorenumerous。Simultaneouslyarisecontrastsofminutestructure。Distincttissuestake
theplaceofmatterthathadpreviouslynorecognizableunlikenessesofparts;
andeachofthetissuesfirstproducedundergoessecondarymodifications,
causingsub—speciesoftissues。Thegranularprotoplasmofthevegetalgerm,
equallywiththatwhichformstheunfoldingpointofeveryshoot,givesorigin
tocellsthatareatfirstalike。Someofthese,astheygrow,flattenand
unitebytheiredgestoformtheouterlayer。Otherslengthen,andatthe
sametimejointogetherinbundlestolaythefoundationofwoody—fibre。
Beforemuchelongating,certainofthesecellsshowabreaking—upofthe
liningdeposit,which,duringelongation,becomesaspiralthread,orareticulated
framework,oraseriesofrings;andbythelongitudinalunionofcellsso
lined,vesselsareformed。Meanwhileeachofthesedifferentiatedtissues
isre—differentiated:instancethatconstitutingtheessentialpartofa
leaf,theupperstratumofwhichiscomposedofchlorophyll—cellsremaining
closelypacked,whilethelowerstratumbecomesspongy。Ofthesamegeneral
characterarethetransformationsundergonebythefertilizedovum,which,
atfirstaclusterofsimilarcells,quicklyreachesastagemarkedbydissimilarity
ofthecells。Morefrequentlyrecurringfissionofthesuperficialcells,
aresultingsmallersizeofthem,andsubsequentunionofthemintoanouter
layer,constitutethefirstdifferentiation;andthemiddleareaofthis
layerisrenderedunliketherestbystillmoreactiveprocessesoflike
kind。Bysuchmodificationsuponmodifications,manyandvarious,arisethe
classesandsub—classesoftissueswhich,intricatelycombinedonewithanother,composeorgans。Equallyconformingtothelawarethechangesingeneralshapeandin
theshapesoforgans。Allgermsareatfirstspheresandalllimbsareat
firstbudsormereroundedlumps。Fromthisprimordialuniformityandsimplicity,
theretakeplacedivergences,bothofthewholesandoftheleadingparts,
towardsmultiformityofcontourandtowardscomplexityofcontour。Remove
thecompactly—foldedyoungleavesthatterminateeveryshoot,andthenucleus
isfoundtobeacentralknobbearinglateralknobs,oneofwhichmaygrow
intoeitheraleaf,asepal,apetal,astamen,oracarpel:alltheseeventually
——unlikepartsbeingatfirstalike。Theshootsthemselvesalsodepartfrom
theirprimitiveunityofform;andwhileeachbranchbecomesmoreorless
differentfromtherest,thewholeexposedpartoftheplantbecomesdifferent
fromtheimbeddedpart。So,too,isitwiththeorgansofanimals。Oneof
theArthropoda,forinstance,haslimbsthatwereoriginallyindistinguishable
fromoneanother——composedahomogeneousseries;butbycontinuousdivergences
therehavearisenamongthemunlikenessesofsizeandform,suchaswesee
inthecrabandthelobster。Vertebratecreaturesequallyexemplifythis
truth。Thewingsandlegsofabirdareofsimilarshapeswhentheybud—outfromthesidesoftheembryo。Thusineveryplantandanimal,conspicuoussecondaryre—distributions
accompanytheprimaryre—distribution。Afirstdifferencebetweentwoparts;
ineachofthesepartsotherdifferenceswhichpresentlybecomeasmarked
asthefirst;andalikemultiplicationofdifferencesingeometricalprogression,
untilthereisreachedthatcomplexcombinationconstitutingtheadult。This
isthehistoryofalllivingthingswhatsoever。PursuinganideawhichHarvey
setafloat,ithasbeenshownbyWolffandVonBaer,thatduringitsdevelopment
eachorganismpassesfromastateofhomogeneitytoastateofheterogeneity。Foragenerationthistruthhasbeenacceptedbybiologists。*