Itisapunylittleinsect,almostastinyasthemidgeswhomweseeeddyinginarayofsunshineattheendofautumn。Itsdressisgoldenbronze;itseyesarecoralred。Itcarriesanakedsword,thatistosay,thesheathofitsdrillstandsoutslantwiseatthetipofitsbelly,insteadoflyinginahollowgroovealongtheback,asitdoeswiththeLeucospis。Thisscabbardholdsthelatterhalfoftheinoculatingfilament,whichextendsbelowtheanimaltothebaseoftheabdomen。Inshort,itsutensilisthatoftheLeucospis,withthisdifference,thatitslowerhalfsticksoutlikearapier。
Thismitethatbearsasworduponherrumpisyetanotherpersecutorofthemasonbeesandnotoneoftheleastformidable。
SheexploitstheirnestsatthesametimeastheLeucospis。Iseeher,liketheLeucospis,slowlyexplorethegroundwithherantennae;Iseeher,liketheLeucospis,bravelydriveherdaggerintothestonewall。Moretakenupwithherwork,lessconsciousperhapsofdanger,shepaysnoheedtothemanwhoisobservinghersoclosely。WheretheLeucospisflies,shedoesnotbudge。Sogreatisherassurancethatshecomesrightintomystudy,tomyworktable,anddisputesmyownershipofthenestswhoseoccupantsIamexamining。Sheoperatesundermylens,sheoperatesjustbesidemyforceps。Whatriskdoessherun?Whatcanonedotoathingsoverysmall?SheissocertainofhersafetythatIcantaketheMason’snestinmyhand,moveit,putitdownandtakeitupagainwithouttheinsect’sraisinganyobjection:itcontinuesitsworkevenwhenmymagnifyingglassisplacedoverit。
OneoftheseheroineshascometoinspectanestoftheChalicodomaoftheWalls,mostofwhosecellsareoccupiedbythenumerouscocoonsofaparasite,theStelis。Thecontentsofthesecells,whichhavebeenpartiallyrippeduptosatisfymycuriosity,areverymuchexposedtoview。Thewindfallappearstobeappreciated,forIseethedwarfferretaboutfromcelltocellforfourdaysonend,seeherchoosehercocoonandinsertherawlinthemostapprovedfashion。Ithuslearnthatsight,althoughanindispensableguideinsearching,doesnotdecideupontheproperspotfortheoperation。Hereisaninsectexploringnotthestonyexteriorofthemason’sdwelling,butthesurfaceofcocoonswovenofsilk。Theexplorerhasneverfoundherselfplacedinsuchcircumstances,norhasanyofherracebeforeher,everycocoon,undernormalconditions,beingprotectedbyasurroundingwall。Nomatter:despitetheprofounddifferenceinthesurfaces,theinsectdoesnotwaver。Warnedbyaspecialsense,anundecipherableriddletoourselves,itknowsthattheobjectofitssearchlieshiddenunderthisunfamiliarcasing。
Thesenseofsmellhasalreadybeenshowntobeoutofthequestion;thatofsightisnoweliminatedinitsturn。
ThatsheshouldborethroughthecocoonsoftheStelis,aparasiteofthemasonbee,doesnotsurprisemeatall:Iknowhowindifferentmyboldvisitoristothenatureofthevictualsdestinedforherfamily。Ihavenoticedherpresenceinthehomesofbeesdifferinggreatlyinsizeandhabits:Anthophorae,Osmiae,Chalicodomae,Anthidia。TheStelisexploitedonmytableisonevictimmore;andthatisall。Theinterestdoesnotliethere。
Theinterestliesinthemaneuversoftheinsect,whichIamabletofollowunderthemostfavorableconditions。
Bentsharplyatrightangles,likeacoupleofbrokenmatches,theantennaefeelthecocoonwiththeirtipsalone。Theterminaljointisthehomeofthisstrangesensewhichdiscernsfromafarwhatnoeyesees,noscentdistinguishesandnoearhears。Ifthepointexploredbefoundsuitable,theinsecthoistsitselfontiptoesoastogivefullscopetotheplayofitsmechanism;itbringsthetipofthebellyalittleforward;andtheentireovipositor——
inoculating—needleandscabbard——standsperpendiculartothecocoon,inthecenterofthequadrilateraldescribedbythefourhindlegs,aneminentlyfavorablepositionforobtainingthemaximumeffect。Forsometime,thewholeoftheawlbearsonthecocoon,feelingallroundwithitspoint,gropingabout;then,suddenly,theboringneedleisreleasedfromitssheath,whichfallsbackalongthebody,whiletheneedlestrivestomakeitsentrance。Theoperationisadifficultone。Iseetheinsectmakeascoreofattempts,oneaftertheother,withoutsucceedinginpiercingthetoughwrapperoftheStelis。Shouldtheinstrumentnotpenetrate,itretreatsintoitssheathandtheinsectresumesitsscrutinyofthecocoon,soundingitpointbypointwiththetipsofitsantennae。Thenfurtherthrustsaretrieduntilonesucceeds。
Theeggsarelittlespindles,whiteandgleaminglikeivory,abouttwo—thirdsofamillimeterinlength。Theyhavenotthelong,curvedpeduncleoftheLeucospis’eggs;theyarenotsuspendedfromtheceilingofthecocoonlikethese,butarelaidwithoutorderaroundthefosteringlarva。Lastly,inasinglecellandwithasinglemother,thereisalwaysmorethanonelaying;andthenumberofeggsvariesconsiderablyineach。TheLeucospis,becauseofhergreatsize,whichrivalsthatofhervictim,theBee,findsineachcellprovisionsenoughforoneandonealone。When,therefore,thereismorethanonesetofeggsinanyonecell,thisisduetoamistakeonherpartandnotapremeditatedresult。Wherethewholerationisrequiredforthemealsofasinglegrub,shewouldtakegoodcarenottoinstallseveralifshecouldhelpit。Hercompetitorisnotcalledupontoobservethesamediscretion。A
Chalicodomagrubgivesthedwarfthewherewithaltoportionascoreofherlittleones,whowillliveincommonandinallcomfortonwhatasinglesonofthegiantesswouldeatupbyhimself。Thetinyboringengineer,therefore,alwayssettlesanumerousfamilyatthesamebanquet。Thebowl,ampleforadozenortwo,isemptiedinperfectharmony。
Curiositymademecountthebrood,toseeifthemotherwasabletoestimatethevictualsandtoproportionthenumberofgueststothesumptuousnessofthefareprovided。Mynotesmentionfifty—fourlarvaeinthecellofamaskedAnthophora(Anthophorapersonata)。
Noothercensusattainedthisfigure。Possibly,twodifferentmothershadlaidtheireggsinthiscrowdedhabitation。WiththeMasonbeeoftheWalls,Iseethenumberoflarvaevary,indifferentcells,betweenfourandtwenty—six;withthemasonbeeoftheSheds,betweenfiveandthirty—six;withthethree—hornedOsmia,whosuppliedmewiththelargestnumberofrecords,betweensevenandtwenty—five;withtheblueOsmia(Osmiacyanea,KIRB。),betweenfiveandsix;withtheStelis(Stelisnasuta),betweenfourandtwelve。
Thefirstreturnandthelasttwoseemtopointtosomerelationbetweentheabundanceofprovisionsandthenumberofconsumers。
WhenthemothercomesuponthebountifullarvaofthemaskedAnthophora,shegivesithalf—a—hundredtofeed;withtheStelisandtheblueOsmia,niggardlyrationsboth,shecontentsherselfwithhalf—a—dozen。Tointroduceintothediningroomonlythenumberofboardersthatthebilloffarewillallowwouldcertainlybeamostdeservingperformance,especiallyastheinsectisplacedunderverydifficultconditionstojudgethecontentsofthecell。
Thesecontents,whichliehiddenundertheceiling,areinvisible;
andtheinsectcanderiveitsinformationonlyfromtheoutsideofthenest,whichvariesinthedifferentspecies。Weshouldthereforehavetoadmittheexistenceofaparticularpowerofdiscrimination,asortofdiscernmentofthespecies,whichisrecognizedaslargeorsmallfromtheoutwardaspectofitshouse。Irefusetogotothislengthinmyconjectures,notthatinstinctseemstomeincapableofsuchfeats,butbecauseoftheparticularsobtainedfromthethree—hornedOsmiaandthetwomasonbees。
Inthecellsofthesethreespecies,IseethenumberoflarvaeputouttonursevaryinsoelasticafashionthatImustabandonallideaofproportionateadjustment。Themother,withouttroublingundulywhethertherebeanexcessoradearthofprovisionsforherfamily,hasfilledthecellsasherfancyprompted,orratheraccordingtothenumberofripeovulescontainedinherovariesatthetimeofthelaying。Iffoodbeover—plentiful,thebroodwillbeallthebetterforitandwillgrowbiggerandstronger;iffoodbescarce,thefamishedyoungsterswillnotdie,butwillremainsmaller。Indeed,withboththelarvaandthefullgrowninsect,I
haveoftenobservedadifferenceinsizewhichvariesaccordingtothedensityofthepopulation,themembersofasmallcolonybeingdoublethesizeoftheirovercrowdedneighbors。
Thegrubsarewhite,taperingatbothends,sharplysegmentedandcoveredallovertheirbodieswithacoatoffine,softhairswhichisinvisibleexceptunderthelens。Theheadconsistsofalittleknobmuchsmallerindiameterthanthebody。Inthishead,themicroscoperevealsmandiblesconsistingoffinespikesofatawnyred,whichspreadintoawide,colorlessbase。Deprivedofanyindentation,incapableofchewinganythingbetweentheirawl—shapedends,thesetwotoolsserveatbesttofixthegrubslightlyatsomepointofthefosteringlarva。Uselessforcarving,therefore,themouthisapureosculatorysucker,whichdrainstheprovisionsbyaprocessofexudationthroughtheskin。WeseehererepeatedwhattheAnthraxandtheLeucospishavealreadyshownus:thegradualexhaustionofavictimwhichtheparasiteconsumeswithoutkillingit。
ItisacuriousspectacleevenafterthatoftheAnthrax。Wehaveheretwentyorthirtystarvelings,allwiththeirmouthspressed,asforakiss,tothebodyoftheplumplarva,which,fromdaytoday,fadesandshrinkswithouttheleastappreciablewound,thuskeepingfreshuntilreducedtoashriveledslough。IfIdisturbthegluttonousswarm,all,withasuddenrecoil,letgo,dropoffandflounderaroundthefostermother。Theyarenolesspromptinresumingtheirsavagekisses。Ineednotaddthatneitheratthepointwheretheyleaveoffnoratthepointwheretheyrecommenceistherethefaintesttraceofliquid。Theoilyexudationoccursonlywhenthepumpisatwork。TolingeroverthisstrangemethodoffeedingissuperfluousafterwhatIhavesaidabouttheAnthrax。
Theappearanceofthefullgrowninsecttakesplaceatthebeginningofsummer,afternearlyawholeyear’sstayintheinvadeddwelling。Thelargenumberofinhabitantsofoneandthesamecellledmetothinkthattheworkofdeliveranceoughttopresentacertaininterest。Theyareallequallyanxioustoclearthewallsoftheprisonattheearliestpossiblemomentandtocomeforthintothegreatfestivalofthesun:dotheyallatthesametime,inaconfusedhorde,attacktheceilingwhichhastobepierced?Istheworkofdeliverancearrangedinthegeneralinterest?Orisindividualselfishnesstheonlyrule?Thesearethequestionswhichobservationwillanswer。
Alittleinadvanceoftheproperseason,Itransfereachfamilyintoashortglasstube,whichwillrepresentthenatalcell。A
good,thickcork,quiteacentimeterdeep,istheobstacletobepiercedforanoutlet。Well,insteadofthemadhasteandtheruinouslackoforganizationwhichIexpectedtofind,mybroodsshowmeintheirglassprisonanexceedinglywellregulatedworkshop。Oneinsect,oneonly,worksatperforatingthecork。
Patiently,withitsmandibles,grainbygrain,itdigsatunnelthewidthofitsbody。Thegalleryissonarrowthat,inordertoreturntothetube,theworkerhastomovebackwards。Itisaslowprocess;andittakeshoursandhourstodigthehole,ahardjobforthefrailminer。
Shouldherfatiguebecometoogreat,theexcavatorleavestheforefrontandmingleswiththecrowd,topolishanddustherself。
Another,thefirstneighborathand,atoncetakesherplaceandisherselfrelievedbyathirdwhenhertaskisdone。Othersagaintaketheirturn,alwaysoneatatime,somuchsothattheworksareneveratastandstillandneverovercrowded。Meanwhile,themultitudekeepsoutoftheway,quietlyandpatiently。Thereisnoanxietyastothedeliverance。Successwillcome:ofthattheyareallconvinced。Whilewaiting,onewashesherantennaebypassingthemthroughhermouth,anotherpolishesherwingswithherhindlegs,anotherfrisksabouttowhileawaytheperiodofinaction。
Somearemakinglove,asovranmeansofkillingtime,whetheronebebornthatdayortwentyyearsago。
Some,Isaid,makelove。Thesefavoredonesarerare;theyhardlycount。Isitthroughindifference?No,butthegallantsarelacking。Thesexesareveryunequallyrepresentedinthepopulationofacell:themalesareinawretchedminorityandsometimesevencompletelyabsent。Thispovertydidnotescapetheolderobservers。Brulle[GaspardAugustBru11e(1809—1873)],theauthorofmanyworksonnaturalhistoryandoneofthefoundersoftheSocieteentomologiquedeFrance),theonlyauthorwhomIamabletoconsultinmyhermitage,says,literally:’Themalesdonotappeartobeknown。’
I,formypart,knowthem;but,consideringtheirfeeblenumber,I
keepaskingmyselfwhatparttheyplayinaharemsodisproportionatetotheirforces。Afewfigureswillshowuswhatmyhesitationsarebasedupon。
Intwenty—twoOsmiacocoons(Osmiatricornis),thetotalcensusoftheinmatesyieldsthreehundredandfifty—four,ofwhomforty—
sevenaremalesandthreehundredandsevenfemales。Theaveragenumberofinmates,therefore,issixteenindividuals;andtherearesixfemalesatleasttoonemale。Thisdisparityismaintained,inmoreorlessmarkedproportions,whateverthespeciesofthebeeinvaded。InthecocoonsoftheMasonbeeoftheSheds,Idiscovertheaverageproportiontobesixfemalestoonemale;inthoseoftheMasonbeeoftheWalls,Ifindonemaletofifteenfemales。
Thesefacts,whichIamunabletostatewithanygreaterprecision,areenoughtogiverisetothesuspicionthatthemales,whoareeventinierdwarfsthanthefemalesandwho,moreover,likeallinsects,areinjuredbyasingleactofpairing,must,inmostcases,remainstrangerstothefemales。Canthemothers,infact,dispensewiththeirassistance,withoutbeingdeprivedofoffspringonthataccount?Idonotsayyes,butIdonotsayno。Thedualityofthesexesisahardproblem。Whytwosexes?Whynotjustone?Itwouldhavebeenmuchsimplerandsavedagreatdealoffoolery。Whysuchathingassex,whenthetuberoftheJerusalemartichokecandowithoutit?Thesearethepregnantquestionssuggestedtome,intheend,byMonodontomeruscupreus,theinsectsoinfinitesimalinbodyandsooverpoweringinnamethatIhadreallyvowednevertospeakofitagainbyitsofficialdesignation。
CHAPTERIVLARVALDIMORPHISM
IfthereaderhaspaidanyattentiontothestoryoftheAnthrax,hemusthaveperceivedthatmynarrativeisincomplete。Thefoxinthefablesawhowthelion’svisitorsenteredhisden,butdidnotseehowtheywentout。Withus,itistheconverse:weknowthewayoutofthemasonbee’sfortress,butwedonotknowthewayin。
Toleavethecellofwhichhehaseatentheowner,theAnthraxbecomesaperforatingmachine,alivingtoolfromwhichourownindustrymighttakeahintifitrequirednewdrillsforboringrocks。Whentheexittunnelisopened,thistoolsplitslikeapodburstinginthesun;andfromthestoutframeworkthereescapesadaintyfly,avelvetyflake,asoftfluffthatastoundsusbyitscontrastwiththeroughnessofthedepthswhenceitascends。Onthispoint,weknowprettywellwhatthereistoknow。Thereremainstheentranceintothecell,apuzzlethathaskeptmeonthealertforaquarterofacentury。
Tobeginwith,itisevidentthatthemothercannotlodgeheregginthecellofthemasonbee,whichhasbeenlongclosedandbarricadedwithacementwallbythetimethattheAnthraxmakesherappearance。Topenetrateit,shewouldhavetobecomeanexcavatingtooloncemoreandresumethecast—offragswhichsheleftbehindintheexitwindow;shewouldhavetoretracehersteps,toberebornapupa;andlifeknowsnoneoftheseretrogressions。Thefullgrowninsect,ifendowedwithclaws,mandiblesandplentyofperseverance,mightatapinchforcethemortarcasket;buttheflyisnotsoendowed。Herslenderlegswouldbestrainedanddeformedbymerelysweepingawayalittledust;hermouthisasuckerforgatheringthesugaryexudationsoftheflowersandnotthesolidpincersneededforthecrumblingofcement。Thereisnoaugereither,noborecopiedfromthatoftheLeucospis,noimplementofanykindthatcanworkitswayintothethicknessofthewallanddispatchtheeggtoitsdestination。Inshort,themotherisabsolutelyincapableofsettlinghereggsinthechamberoftheMasonbee。
Canitbethegrubthatmakesitsownwayintothestoreroom,thatsamegrubwhichwehaveseendrainingtheChalicodomawithitsleech—likekisses?Letuscallthecreaturetomind:alittleoilysausage,whichstretchesandcurlsupjustwhereitlies,withoutbeingabletoshiftitsposition。Itsbodyisasmoothcylinder;
itsmouthsimplyacircularlip。Notoneambulatoryorgandoesitpossess;notevenhairs,protuberancesorwrinklestoenableittocrawl。Theanimalismadefordigestionandimmobility。Itsorganizationisincompatiblewithmovement;everythingtellsussointheclearestfashion。No,thisgrubisevenlessablethanthemothertomakeitswayunaidedintothemason’sdwelling。Andyettheprovisionsarethere;thoseprovisionsmustbereached:itisamatteroflifeordeath;tobeornottobe。Thenhowdoestheflysetaboutit?Itwouldbevainformetoquestionprobabilities,toooftenillusory;toobtainareplyofanyvalue,Ihavebutoneresource;ImustattemptthenearlyimpossibleandwatchtheAnthraxfromtheeggonwards。
AlthoughAnthraxfliesarefairlycommon,inthesenseoftherebeingseveraldifferentspecies,theyarenotplentifulwhenitisacaseofwantingacolonypopulousenoughtoadmitofcontinuousobservation。Iseethem,nowhere,nowthere,inthefiercelysun—
scorchedplaces,flittinghitherandthitherontheoldwalls,theslopesandthesand,sometimesinsmallplatoons,mostoftensingly。Icanexpectnothingofthosevagabonds,whoareheretodayandgonetomorrow,forIknownothingoftheirsettlements。
Tokeepawatchonthem,onebyone,intheblazingheat,isverypainfulandveryunfruitful,astheswift—wingedinsecthasahabitofdisappearingoneknowsnotwhitherjustwhenaprospectofcapturingitssecretbeginstooffer。Ihavewastedmanyapatienthouratthispursuit,withouttheleastresult。
TheremightbesomechanceofsuccesswithAnthraxflieswhosehomewasknowntousbeforehand,especiallyifinsectsofthesamespeciesformedaprettynumerouscolony。Theinquiriesbegunwithonewouldbecontinuedwithasecondandwithmore,untilacompleteverdictwasforthcoming。Now,inthecourseofmylongentomologicalcareer,IhavemetwithbuttwospeciesofAnthraxthatfulfilledthisconditionandweretobefoundregularly:oneatCarpentras;theotheratSerignan。Thefirst,Anthraxsinuata,FALLEN,livesinthecocoonsofOsmiatricornis,whoherselfbuildshernestintheoldgalleriesofthehairy—footedAnthophora;thesecond,Anthraxtrifasciata,MEIGEN,exploitstheChalicodomaoftheSheds。Iwillconsultboth。
Oncemore,hereamI,somewhatlateinlife,atCarpentras,whoserudeGallicnamesetsthefoolsmilingandthescholarthinking。
DearlittletownwhereIspentmytwentiethyearandleftthefirstbitsofmyfleeceuponlife’sbushes,myvisitoftodayisapilgrimage;Ihavecometolaymyeyesoncemoreupontheplacewhichsawthebirthoftheliveliestimpressionsofmyearlydays。
Ibow,inpassing,totheoldcollegewhereItriedmyprenticehandasateacher。Itsappearanceisunchanged;itstilllookslikeapenitentiary。Thoseweretheviewsofourmediaevaleducationalsystem。Tothegaietyandactivityofboyhood,whichwereconsideredunwholesome,itappliedtheremedyofnarrowness,melancholyandgloom。Itshousesofinstructionwere,aboveall,housesofcorrection。ThefreshnessofVirgilwasinterpretedinthestiflingatmosphereofaprison。Icatchaglimpseofayardbetweenfourhighwalls,asortofbearpit,wherethescholarsfoughtforroomfortheirgamesunderthespreadingbranchesofaplanetree。Allaroundwerecellsthatlookedlikehorseboxes,withoutlightorair;thoseweretheclassrooms。Ispeakinthepasttense,fordoubtlessthepresentdayhasseenthelastofthisacademicdestitution。
Hereisthetobaccoshopwhere,onWednesdayevening,comingoutofthecollege,Iwouldbuyoncreditthewherewithaltofillmypipeandthustocelebrateontheevethejoysofthemorrow,thatblessedThursday[theweeklyhalf—holidayinFrenchschools]whichIconsideredsowellemployedinsolvinghardequations,experimentingwithnewchemicalreagents,collectingandidentifyingmyplants。Iwouldmakemytimidrequest,pretendingtohavecomeoutwithoutmymoney,foritishardforaself—
respectingmantoadmitthatheispenniless。Mycandorappearstohaveinspiredsomelittleconfidence;andIobtainedcredit,anunprecedentedthing,withtherepresentativeoftherevenue。[ThegovernmentinFrancehasthesolecontrolofthetobaccotrade,whichformsanimportantbranchoftheinlandrevenue。]Ah,whydidnotIopenashopandexposeforsalesomepacketsofcandles,adozendriedcod,abarrelofsardinesandafewcakesofsoap!Iamnomoreofafoolnoranylessindustriousthananother;andI
shouldhavemademyway。But,asitwas,whatcouldIexpect?Asanaccoucheurofbrains,amolderofintellects,Ihadnoclaimeventobreadandcheese。
Hereismyformerhabitation,occupiedsincebydroningmonks。Intheembrasureofthatwindow,shelteredfromprofanehands,betweentheclosedoutershuttersandthepanes,Iusedtokeepmychemicals,boughtforafewsouscheatedoutoftheweeklybudgetintheearlydaysofourhousekeeping。Thebowlofapipewasmycrucible,asweetjarmyretort,mustardpotsmyreceptaclesforoxidesandsulfides。Myexperiments,harmlessordangerous,weremadeonacornerofthefirebesidethesimmeringbroth。
HowIshouldlovetoseethatroomagainwhereIporedoverdifferentialsandintegrals,whereIcalmedmypoorburningheadbygazingatMontVentoux,whosesummitheldinstoreformycomingexpedition’thosedenizensofarcticclimes,thesaxifrageandthepoppy!Andtoseemyfamiliarfriend,theblackboardwhichIhiredatfivefrancsayearfromacrustyjoiner,thatboardwhosevalueIpaidmanytimesover,thoughI。couldneverbuyitoutright,forwantofthenecessarycash!TheconicsectionswhichIdescribedonthatblackboard,thelearnedhieroglyphics!
Thoughallmyefforts,whichwerethemoredeservingbecauseIhadtoworkalone,ledtoalmostnothinginthatcongenialcalling,I
wouldbeginitalloveragainifIcould。IshouldlovetobeconversingforthefirsttimewithLeibnitzandNewton,withLaplaceandLagrange,withCuvierandJussieu,evenifIhadafterwardstosolvethatotherarduousproblem:howtoprocureone’sdailybread。Ah,youngmen,mysuccessors,whataneasytimeyouhaveofittoday!Ifyoudon’tknowit,thenletmetellyousobymeansofthesefewpagesfromthelifeofoneofyourelders。
Butletusnotforgetourinsects,whilelisteningtotheechoesofillusionsanddifficultiesrousedinmymemoriesbythecupboardwindowandthehiredblackboard。LetusgobacktothesunkenroadsoftheLegue,whichhavebecomeclassic,sotheysay,sincetheappearanceofmynotesontheOilbeetles。Yeillustriousravines,withyoursun—bakedslopes,ifIhavecontributedalittletoyourfame,you,inyourturn,havegivenmemanyfairhoursofforgetfulnessinthehappinessoflearning。You,atleast,didnotluremewithvainhopes;allthatyoupromisedyougavemeandoftenahundredfold。Youaremypromisedland,whereIwouldhavesoughtatthelasttopitchmyobserver’stent。Mywishwasnottoberealized。Letme,atleast,inpassing,greetmybelovedanimalsoftheolddays。
IraisemyhattoCerceristuberculata,whomIseeengagedonthatslant,storingherCleonus[alargespeciesofweevil]。AsIsawherthen,soIseehernow:thesamestaggeringattemptstohoistthepreytothemouthoftheburrow;thesamebrawlsbetweenmaleswatchinginthebrushwoodofthekermesoak。Thesightofthemsendsayoungerbloodcoursingthroughmyveins;Ireceiveasitwerethebreathofanewspringtimeoflife。Timepresses;letuspasson。
Anotherbowonthisside。Ihearbuzzingupabove,onthatledge,acolonyofSphexwasps,stabbingtheircrickets。Wewillgivethemafriendlyglance,butnomore。Myacquaintancesherearetoonumerous;Ihavenottheleisuretorenewmyformerrelationswithallofthem。Withoutstopping,awaveofthehattothePhilanthi[bee—huntingwasps]whosendthelongavalanchesofrubbishstreamingdownfromtheirnests;andtoStizusruficornis,[ahuntingwasp]whostacksherprayingmantisesbetweentwoflakesofsandstone;andtothesilkyAmmophila[adiggerwasp]withtheredlegs,whocollectsanundergroundstoreofloopers[alsoknownasmeasuringworms,thelarvaeorcaterpillarsofthegeometridmoth]
andtotheTachtyti[huntingwasps],devourersoflocusts;andtotheEumenes,buildersofclaycupolasonabough。
Hereweareatlast。Thishigh,perpendicularrock,facingthesouthtoalengthofsomehundredsofyardsandriddledwithholeslikeamonstroussponge,isthetime—honoreddwellingplaceofthehairy—footedAnthophoraandofherrentfreetenant,thethree—
hornedOsmia。Herealsoswarmtheirexterminators:theSitarisbeetle,theparasiteoftheAnthophora;theAnthraxfly,themurdereroftheOsmia。Illinformedastotheproperperiod,I
havecomeratherlate,onthe10thofSeptember。Ishouldhavebeenhereamonthago,orevenbytheendofJuly,towatchthefly’soperations。Myjourneythreatenstobefruitless:IseebutafewrareAnthraxflies,hoveringroundthefaceofthecliff。Wewillnotdespair,however,andwewillbeginbyconsultingthelocality。
TheAnthophora’scellscontainthisbeeinthelarvalstage。SomeofthemprovidemewiththeoilbeetleandtheSitaris,rarefindsatonetime,todayofnousetome。OtherscontaintheMelecta[aparasiticbee]intheformofahighlycoloredpupa,oreveninthatofthefullgrowninsect。TheOsmia,stillmoreprecocious,thoughdatingfromthesameperiod,showsherselfexclusivelyintheadultform,abadomenformyinvestigations,forwhattheAnthraxdemandsisthelarvaandnottheperfectinsect。Thefly’sgrubdoublesmyapprehensions。Itsdevelopmentiscomplete,thelarvaonwhichitfeedsisconsumed,perhapsseveralweeksago。I
nolongerdoubtbutthatIhavecometoolatetoseewhathappensintheOsmia’scocoons。
Isthegamelost?Notyet。MynotescontainevidenceofAnthraxflieshatchinginthelatterhalfofSeptember。Besides,thosewhomInowseeexploringtherockarenottheretotakeexercise:
theirpreoccupationisthesettlingofthefamily。ThesebelatedonescannottackletheOsmia,who,withherfirm,adultflesh,wouldnotsuitthenursling’sdelicateneedsandwho,moreover,powerfulassheis,wouldofferresistance。Butinautumnalessnumerouscolonyofhoneygathererstakestheplace,upontheslope,ofthespringcolony,fromwhichitdiffersinspecies。Inparticular,IseetheDiademAnthidium[aclothierbeewholineshernestwithwoolandcotton]atwork,enteringhergalleriesatonetimewithherharvestofpollendustandatanotherwithherlittlebaleofcotton。MightnottheseautumnalBeesbethemselvesexploitedbytheAnthrax,thesamethatselectedtheOsmiaashervictimacoupleofmonthsearlier?ThiswouldexplainthepresenceoftheAnthraxflieswhomInowseefussingabout。
Alittlereassuredbythisconjecture,Itakemystandatthefootoftherock,underabroilingsun;and,forhalfaday,Ifollowtheevolutionsofmyflies。Theyflitquietlyinfrontoftheslope,atafewinchesfromtheearthycovering。Theygofromoneorificetothenext,butwithoutevenpenetrating。Forthatmatter,theirbigwings,extendedcrosswiseevenwhenatrest,wouldresisttheirentranceintoagallery,whichistoonarrowtoadmitthosespreadingsails。Andsotheyexplorethecliff,goingtoandfroandupanddown,withaflightthatisnowsudden,nowsmoothandslow。Fromtimetotime,IseetheAnthraxquicklyapproachthewallandlowerherabdomenasthoughtotouchtheearthwiththeendofherovipositor。Thisproceedingtakesnolongerthanthetwinklingofaneye。Whenitisdone,theinsectalightselsewhereandrests。Thenitresumesitssoberflight,itslonginvestigationsanditssuddenblowswiththetipofitsbellyagainstthelayerofearth。TheBombylii[beeflies]
observesimilartacticswhensoaringatashortheightabovetheground。
Iatoncerushedtothespottouched,lensinhand,inthehopeoffindingtheeggwhicheverythingtoldmewaslaidduringthattapoftheabdomen。Icoulddistinguishnothing,inspiteoftheclosestattention。Itistruethatmyexhaustion,togetherwiththeblindinglightandscorchingheat,madeexaminationverydifficult。Afterwards,whenImadetheacquaintanceofthetinythingthatissuesfromthategg,myfailurenolongersurprisedme。
Intheleisureofmystudy,withmyeyesrestedandwithmymostpowerfulglassesheldinahandnolongershakingwithexcitementandfatigue,Ihavetheverygreatestdifficultyinfindingtheinfinitesimalcreature,thoughIknowexactlywhereitlies。ThenhowcouldIseetheegg,wornoutasIwasunderthesun—bakedcliff,howdiscovertheprecisespotofalayingperformedinamomentbyaninsectseenonlyatadistance?InthepainfulconditionswhereinIfoundmyself,failurewasinevitable。
Despitemynegativeattempts,therefore,IremainconvincedthattheAnthraxfliesstrewtheireggsonebyone,onthespotsfrequentedbythosebeeswhosuittheirgrubs。Eachoftheirsuddenstrokeswiththetipoftheabdomenrepresentsalaying。
Theytakenoprecautiontoplacethegermundercover;forthatmatter,anysuchprecautionwouldberenderedimpossiblebythemother’sstructure。Theegg,thatdelicateobject,islaidroughlyintheblazingsun,betweengrainsofsand,insomewrinkleofthecalcinedchalk。Thatsummaryinstallationissufficient,providedthecovetedlarvabenearathand。Itisfortheyounggrubnowtomanageasbestitcanatitsownriskandperil。
ThoughthesunkenroadsoftheLeguedidnottellmeallthatI
wishedtoknow,theyatleastmadeitveryprobablethatthecominggrubmustreachthevictualledcellbyitsownefforts。Butthegrubwhichweknow,theonethatdrainsthebagoffatwhichmaybeaChalicodomalarvaoranOsmialarva,cannotmovefromitsplace,stilllessindulgeinjourneysofdiscoverythroughthethicknessofawallandthewebofacocoon。Soanimperativenecessitypresentsitself:theremustperforcebeaninitiallarvaform,capableofmovingandorganizedforsearching,aformunderwhichthegrubwouldattainitsend。TheAnthraxwouldthuspossesstwolarvalstates:onetopenetratetotheprovisions;theothertoconsumethem。Iallowmyselftobeconvincedbythelogicofitall;Ialreadyseeinmymind’seyetheweeanimalcomingoutoftheegg,endowedwithsufficientpowerofmotionnottodreadawalkandwithsufficientslendernesstoglideintothesmallestcrevices。Onceinthepresenceofthelarvaonwhichitistofeed,itdoffsitstravellingdressandbecomestheobeseanimalwhoseonedutyitistogrowbigandfatinimmobility。Thisisallverycoherent;itisalldeducedlikeageometricalproposition。Buttothewingsofimagination,howeversmooththeirflight,wemustpreferthesandalsofobservedfacts,theslowsandalswiththeleadensoles。Thusshod,Iproceed。
Nextyear,Iresumemyinvestigations,thistimeontheAnthraxoftheChalicodoma,whoismyneighborinthesurroundingwastelandsandwillallowmetorepeatmyvisitsdaily,morningandeveningifneedbe。Taughtbymyearlierstudies,InowknowtheexactperiodoftheBee’shatchingandthereforeoftheAnthrax’laying,whichmusttakeplacesoonafter。AnthraxtrifasciatasettlesherfamilyinJuly,orinAugustatlatest。Everymorning,atnineo’clock,whentheheatbeginstobeunendurableandwhen,touse[theauthor’sgardenerandfactotum]Favier’sexpression,anextralogisflungonthebonfireofthesun,Itakethefield,preparedtocomebackwithmyheadachingfromtheglare,providedthatIbringhomethesolutionofmypuzzle。Amanmusthavethedevilinhimtoleavetheshadeatthistimeoftheyear。Andwhatfor,pray?