CHARLESDARWINTOJ。D。HOOKER。
  Down,June10[1864]……IhavenowreadtwoGermanbooks,andallIbelievethathasbeenwrittenonclimbers,andithasstirredmeuptofindthatIhaveagooddealofnewmatter。Itisstrange,butIreallythinknoonehasexplainedsimpletwiningplants。Thesebookshavestirredmeup,andmademewishforplantsspecifiedinthem。Ishallbeverygladofthoseyoumention。
  IhavewrittentoVeitchforyoungNepenthesandVanilla(whichIbelievewillturnoutagrandcase,thougharootcreeper),ifIcannotbuyyoungVanillaIwillaskyou。Ihaveorderedaleaf—climbingfern,Lygodium。
  Allthisworkaboutclimberswouldhurtmyconscience,didIthinkIcoulddoharderwork。(Hewasmuchoutofhealthatthistime。)
  [Hecontinuedhisobservationsonclimbingplantsduringtheprolongedillnessfromwhichhesufferedintheautumnof1863,andinthefollowingspring。HewrotetoSirJ。D。Hooker,apparentlyinMarch1864:——
  "ForseveraldaysIhavebeendecidedlybetter,andwhatIlaymuchstresson(whateverdoctorssay),mybrainfeelsfarstronger,andIhavelostmanydreadfulsensations。Thehot—houseissuchanamusementtome,andmyamusementIowetoyou,asmydelightistolookatthemanyoddleavesandplantsfromKew……TheonlyapproachtoworkwhichIcandoistolookattendrilsandclimbers,thisdoesnotdistressmyweakenedbrain。AskOlivertolookovertheenclosedqueries(anddoyoulook)andamuseabroken—downbrothernaturalistbyansweringanywhichhecan。Ifyoueverloungethroughyourhouses,remembermeandclimbingplants。"
  OnOctober29,1864,hewrotetoDr。Gray:——
  "Ihavenotbeenabletoresistdoingalittlemoreatyourgodchild,myclimbingpaper,orratherinsizelittlebook,whichbyJoveIwillhavecopiedout,elseIshallneverstop。Thishasbeennewsortofworkforme,andIhavebeenpleasedtofindwhatacapitalguideforobservationsafullconvictionofthechangeofspeciesis。"
  OnJanuary19,1865,hewrotetoSirJ。D。Hooker:——
  "Itisworkinghours,butIamtryingtotakeaday’sholiday,forI
  finishedanddespatchedyesterdaymyclimbingpaper。ForthelasttendaysIhavedonenothingbutcorrectrefractorysentences,andIloathethewholesubject。"
  AlettertoDr。Gray,April9,1865,hasawordortwoonthesubject:——
  "Ihavebeguncorrectingproofsofmypaperon’ClimbingPlants。’I
  supposeIshallbeabletosendyouacopyinfourorfiveweeks。Ithinkitcontainsagooddealnewandsomecuriouspoints,butitissofearfullylong,thatnoonewilleverreadit。If,however,youdonotSKIMthroughit,youwillbeanunnaturalparent,foritisyourchild。"
  Dr。Graynotonlyreaditbutapprovedofit,tomyfather’sgreatsatisfaction,asthefollowingextractsshow:——
  "IwasmuchpleasedtogetyourletterofJuly24th。NowthatIcandonothing,Imaunderoveroldsubjects,andyourapprobationofmyclimbingpapergivesmeVERYgreatsatisfaction。ImademyobservationswhenI
  coulddonothingelseandmuchenjoyedit,butalwaysdoubtedwhethertheywereworthpublishing。IdemurtoitsnotbeingnecessarytoexplainindetailaboutthespiresinCAUGHTtendrilsrunninginoppositedirections;
  forthefactforalongtimeconfoundedme,andIhavefounditdifficultenoughtoexplainthecausetotwoorthreepersons。"(August15,1865。)
  "Ireceivedyesterdayyourarticle(IntheSeptembernumberof’Silliman’sJournal,’concludedintheJanuarynumber,1866。)onclimbers,andithaspleasedmeinanextraordinaryandevensillymanner。Youpaymeasuperbcompliment,andasIhavejustsaidtomywife,IthinkmyfriendsmustperceivethatIlikepraise,theygivemesuchheartydoses。Ialwaysadmireyourskillinreviewsorabstracts,andyouhavedonethisarticleexcellentlyandgiventhewholeessenceofmypaper……IhavehadaletterfromagoodZoologistinS。Brazil,F。Muller,whohasbeenstirreduptoobserveclimbersandgivesmesomecuriouscasesofBRANCH—climbers,inwhichbranchesareconvertedintotendrils,andthencontinuetogrowandthrowoutleavesandnewbranches,andthenlosetheirtendrilcharacter。"
  (October1865。)
  ThepaperonClimbingPlantswasrepublishedin1875,asaseparatebook。
  Theauthorhadbeenunabletogivehiscustomaryamountofcaretothestyleoftheoriginalessay,owingtothefactthatitwaswrittenduringaperiodofcontinuedill—health,anditwasnowfoundtorequireagreatdealofalteration。HewrotetoSirJ。D。Hooker(March3,1875):"Itisluckyforauthorsingeneralthattheydonotrequiresuchdreadfulworkinmerelylickingwhattheywriteintoshape。"AndtoMr。MurrayinSeptemberhewrote:"Thecorrectionsareheavyin’ClimbingPlants,’andyetI
  deliberatelywentovertheMS。andoldsheetsthreetimes。"ThebookwaspublishedinSeptember1875,aneditionof1500copieswasstruckoff;theeditionsoldfairlywell,and500additionalcopieswereprintedinJuneofthefollowingyear。]
  INSECTIVOROUSPLANTS。
  [Inthesummerof1860hewasstayingatthehouseofhissister—in—law,MissWedgwood,inAshdownForest,whencehewrote(July29,1860),toSirJosephHooker;——
  "LatterlyIhavedonenothinghere;butatfirstIamusedmyselfwithafewobservationsontheinsect—catchingpowerofDrosera;andImustconsultyousometimewhethermy’twaddle’isworthcommunicatingtotheLinneanSociety。"
  InAugusthewrotetothesamefriend:——
  "IwillgratefullysendmynotesonDroserawhencopiedbymycopier:thesubjectamusedmewhenIhadnothingtodo。"
  Hehasdescribedinthe’Autobiography’(volumei。),thegeneralnatureoftheseearlyexperiments。Henoticedinsectsstickingtotheleaves,andfindingthatflies,etc。,placedontheadhesiveglandswereheldfastandembraced,hesuspectedthattheleaveswereadaptedtosupplynitrogenousfoodtotheplant。Hethereforetriedtheeffectontheleavesofvariousnitrogenousfluids——withresultswhich,asfarastheywent,verifiedhissurmise。InSeptember,1860,hewrotetoDr。Gray:——
  "IhavebeeninfinitelyamusedbyworkingatDrosera:themovementsarereallycurious;andthemannerinwhichtheleavesdetectcertainnitrogenouscompoundsismarvellous。Youwilllaugh;butitis,atpresent,myfullbelief(afterendlessexperiments)thattheydetect(andmoveinconsequenceof)the1/2880partofasinglegrainofnitrateofammonia;butthemuriateandsulphateofammoniabothertheirchemicalskill,andtheycannotmakeanythingofthenitrogeninthesesalts!I
  beganthisworkonDroserainrelationtoGRADATIONasthrowinglightonDionaea。"
  LaterintheautumnhewasagainobligedtoleavehomeforEastbourne,wherehecontinuedhisworkonDrosera。Theworkwassonewtohimthathefoundhimselfindifficultiesinthepreparationofsolutions,andbecamepuzzledoverfluidandsolidounces,etc。etc。Toafriend,thelateMr。
  E。Cresy,whocametohishelpinthematterofweightsandmeasures,hewrotegivinganaccountoftheexperiments。Theextract(November2,1860)
  whichfollowsillustratesthealmostsuperstitiousprecautionsheoftenappliedtohisresearches:——
  "GenerallyIhavescrutinisedeveryglandandhairontheleafbeforeexperimenting;butitoccurredtomethatImightinsomewayaffecttheleaf;thoughthisisalmostimpossible,asIscrutinisedwithequalcarethosethatIputintodistilledwater(thesamewaterbeingusedfordissolvingthecarbonateofammonia)。Ithencutofffourleaves(nottouchingthemwithmyfingers),andputtheminplainwater,andfourotherleavesintotheweaksolution,andafterleavingthemforanhourandahalf,Iexaminedeveryhaironalleightleaves;nochangeonthefourinwater;everyglandandhairaffectedinthoseinammonia。
  "Ihadmeasuredthequantityofweaksolution,andIcountedtheglandswhichhadabsorbedtheammonia,andwereplainlyaffected;theresultconvincedmethateachglandcouldnothaveabsorbedmorethan1/64000or1/65000ofagrain。Ihavetriednumbersofotherexperimentsallpointingtothesameresult。Someexperimentsleadmetobelievethatverysensitiveleavesareactedonbymuchsmallerdoses。Reflecthowlittleammoniaaplantcangetgrowingonpoorsoil——yetitisnourished。Thereallysurprisingpartseemstomethattheeffectshouldbevisible,andnotunderveryhighpower;foraftertryingahighpower,Ithoughtitwouldbesafernottoconsideranyeffectwhichwasnotplainlyvisibleunderatwo—thirdsobjectglassandmiddleeye—piece。Theeffectwhichthecarbonateofammoniaproducesisthesegregationofthehomogeneousfluidinthecellsintoacloudofgranulesandcolourlessfluid;andsubsequentlythegranulescoalesceintolargermasses,andforhourshavetheoddestmovements——coalescing,dividing,coalescingadinfinitum。Idonotknowwhetheryouwillcarefortheseill—writtendetails;but,asyouasked,IamsureIamboundtocomply,afteralltheverykindandgreattroublewhichyouhavetaken。"
  OnhisreturnhomehewrotetoSirJ。D。Hooker(November21,1860):——
  "IhavebeenworkinglikeamadmanatDrosera。Hereisafactforyouwhichiscertainasyoustandwhereyouare,thoughyouwon’tbelieveit,thatabitofhair1/78000ofonegraininweightplacedongland,willcauseONEofthegland—bearinghairsofDroseratocurveinwards,andwillaltertheconditionofthecontentsofeverycellinthefoot—stalkofthegland。"
  AndafewdayslatertoLyell:——
  "IwillandmustfinishmyDroseraMS。,whichwilltakemeaweek,for,atthepresentmoment,IcaremoreaboutDroserathantheoriginofallthespeciesintheworld。ButIwillnotpublishonDroseratillnextyear,forIamfrightenedandastoundedatmyresults。Ideclareitisacertainfact,thatoneorganissosensitivetotouch,thataweightseventy—eighttimeslessthanthat,viz。,1/1000ofagrain,whichwillmovethebestchemicalbalance,sufficestocauseaconspicuousmovement。Isitnotcuriousthataplantshouldbefarmoresensitivetothetouchthananynerveinthehumanbody?YetIamperfectlysurethatthisistrue。WhenIamonmyhobby—horse,Inevercanresisttellingmyfriendshowwellmyhobbygoes,soyoumustforgivetherider。"
  Theworkwascontinued,asaholidaytask,atBournemouth,wherehestayedduringtheautumnof1862。Thediscussioninthefollowingletteron"nervousmatter"inDroseraisofinterestinrelationtorecentresearchesonthecontinuityofprotoplasmfromcelltocell:]
  CHARLESDARWINTOJ。D。HOOKER。
  CliffCottage,Bournemouth。
  September26[1862]。
  MydearHooker,Donotreadthistillyouhaveleisure。Ifthatblessedmomentevercomes,Ishouldbeverygladtohaveyouropiniononthesubjectofthisletter。
  IamledtotheopinionthatDroseramusthavediffusedmatterinorganicconnection,closelyanalogoustothenervousmatterofanimals。Whentheglandsofoneofthepapillaeortentacles,initsnaturalpositionissuppliedwithnitrogenisedfluidandcertainotherstimulants,orwhenloadedwithanextremelyslightweight,orwhenstruckseveraltimeswithaneedle,thepedicelbendsnearitsbaseinunderoneminute。Thesevariedstimulantsareconveyeddownthepedicelbysomemeans;itcannotbevibration,fordropsoffluidputonquitequietlycausethemovement;itcannotbeabsorptionofthefluidfromcelltocell,forIcanseetherateofabsorption,whichthoughquick,isfarslower,andinDionaeathetransmissionisinstantaneous;analogyfromanimalswouldpointtotransmissionthroughnervousmatter。Reflectingontherapidpowerofabsorptionintheglands,theextremesensibilityofthewholeorgan,andtheconspicuousmovementcausedbyvariedstimulants,Ihavetriedanumberofsubstanceswhicharenotcausticorcorrosive,……butmostofwhichareknowntohavearemarkableactiononthenervousmatterofanimals。Youwillseetheresultsintheenclosedpaper。Asthenervousmatterofdifferentanimalsaredifferentlyactedonbythesamepoisons,onewouldnotexpectthesameactiononplantsandanimals;onlyifplantshavediffusednervousmatter,somedegreeofanalogousaction。Andthisispartiallythecase。Consideringtheseexperiments,togetherwiththepreviouslymaderemarksonthefunctionsoftheparts,Icannotavoidtheconclusion,thatDroserapossessesmatteratleastinsomedegreeanalogousinconstitutionandfunctiontonervousmatter。Nowdotellmewhatyouthink,asfarasyoucanjudgefrommyabstract;ofcoursemanymoreexperimentswouldhavetobetried;butinformeryearsItriedonthewholeleaf,insteadofonseparateglands,anumberofinnocuous(Thislineofinvestigationmadehimwishforinformationontheactionofpoisonsonplants;asinmanyothercasesheappliedtoProfessorOliver,andinreferencetotheresultwrotetoHooker:"PraythankOliverheartilyforhisheapofreferencesonpoisons。")substances,suchassugar,gum,starch,etc。,andtheyproducednoeffect。Youropinionwillaidmeindecidingsomefutureyearingoingonwiththissubject。Ishouldnothavethoughtitworthattempting,butIhadnothingonearthtodo。
  MydearHooker,Yoursverysincerely,CH。DARWIN。
  P。S。——WereturnhomeonMonday28th。ThankHeaven!
  [Alongbreaknowensuedinhisworkoninsectivorousplants,anditwasnottill1872thatthesubjectseriouslyoccupiedhimagain。ApassageinalettertoDr。AsaGray,writtenin1863or1864,shows,however,thatthequestionwasnotaltogetherabsentfromhismindintheinterim:——
  "DependonityouareunjustonthemeritsofmybelovedDrosera;itisawonderfulplant,orratheramostsagaciousanimal。IwillstickupforDroseratothedayofmydeath。HeavenknowswhetherIshalleverpublishmypileofexperimentsonit。"
  Henotesinhisdiarythatthelastproofofthe’ExpressionoftheEmotions’wasfinishedonAugust22,1872,andthathebegantoworkonDroseraonthefollowingday。]
  CHARLESDARWINTOASAGRAY。
  [Sevenoaks],October22[1872]……IhaveworkedprettyhardforfourorfiveweeksonDrosera,andthenbrokedown;sothatwetookahousenearSevenoaksforthreeweeks(whereI
  nowam)togetcompleterest。Ihaveverylittlepowerofworkingnow,andmustputofftherestoftheworkonDroseratillnextspring,asmyplantsaredying。Itisanendlesssubject,andImustcutitshort,andforthisreasonshallnotdomuchonDionaea。ThepointwhichhasinterestedmemostistracingtheNERVES!whichfollowthevascularbundles。Byaprickwithasharplancetatacertainpoint,Icanparalyseone—halftheleaf,sothatastimulustotheotherhalfcausesnomovement。Itisjustlikedividingthespinalmarrowofafrog:——nostimuluscanbesentfromthebrainoranteriorpartofthespinetothehindlegs;butiftheselatterarestimulated,theymovebyreflexaction。Ifindmyoldresultsabouttheastonishingsensitivenessofthenervoussystem(!?)ofDroseratovariousstimulantsfullyconfirmedandextended……
  [HisworkondigestioninDroseraandotherpointsinthephysiologyoftheplantsoonledhimintoregionswherehisknowledgewasdefective,andheretheadviceandassistancewhichhereceivedfromDr。BurdonSandersonwasofmuchvalue:]
  CHARLESDARWINTOJ。BURDONSANDERSON。
  Down,July25,1873。
  MydearDr。Sanderson,IshouldliketotellyoualittleaboutmyrecentworkwithDrosera,toshowthatIhaveprofitedbyyoursuggestions,andtoaskaquestionortwo。
  1。ItisreallybeautifulhowquicklyandwellDroseraandDionaeadissolvelittlecubesofalbumenandgelatine。Ikeptthesamesizedcubesonwetmossforcomparison。WhenyouwerehereIforgotthatIhadtriedgelatine,butalbumenisfarbetterforwatchingitsdissolutionandabsorption。Franklandhastoldmehowtotestinaroughwayforpepsin;
  andintheautumnhewilldiscoverwhatacidthedigestivejuicecontains。
  2。Adecoctionofcabbage—leavesandgreenpeascausesasmuchinflectionasaninfusionofrawmeat;adecoctionofgrassislesspowerful。ThoughIhearthatthechemiststrytoprecipitateallalbumenfromtheextractofbelladonna,Ithinktheymustfail,astheextractcausesinflection,whereasanewlotofatropine,aswellasthevalerianate[ofatropine],producenoeffect。
  3。Ihavebeentryingagoodmanyexperimentswithheatedwater……Shouldyounotcallthefollowingcaseoneofheatrigor?Twoleaveswereheatedto130deg,andhadeverytentaclecloselyinflected;onewastakenoutandplacedincoldwater,anditre—expanded;theotherwasheatedto145deg,andhadnottheleastpowerofre—expansion。Isnotthislattercaseheatrigor?Ifyoucaninformme,Ishouldverymuchliketohearatwhattemperaturecold—bloodedandinvertebrateanimalsarekilled。
  4。Imusttellyoumyfinalresult,ofwhichIamsure,[asto]thesensitivenessofDrosera。Imadeasolutionofonepartofphosphateofammoniabyweightto218,750ofwater;ofthissolutionIgavesomuchthataleafgot1/8000ofagrainofthephosphate。Ithencountedtheglands,andeachcouldhavegotonly1/1552000ofagrain;thisbeingabsorbedbytheglands,sufficedtocausethetentaclesbearingtheseglandstobendthroughanangleof180deg。Suchsensitivenessrequireshotweather,andcarefullyselectedyoungyetmatureleaves。Itstrikesmeasawonderfulfact。ImustaddthatItookeveryprecaution,bytryingnumerousleavesatthesametimeinthesolutionandinthesamewaterwhichwasusedformakingthesolution。
  5。Ifyoucanpersuadeyourfriendtotrytheeffectsofcarbonateofammoniaontheaggregationofthewhitebloodcorpuscles,Ishouldverymuchliketoheartheresult。
  Ihopethisletterwillnothaveweariedyou。
  Believeme,yoursverysincerely,CHARLESDARWIN。
  CHARLESDARWINTOW。THISELTONDYER。
  Down,24[December1873?]。
  MydearMr。Dyer,Ifearthatyouwillthinkmeagreatbore,butIcannotresisttellingyouthatIhavejustfoundoutthattheleavesofPinguiculapossessabeautifullyadaptedpowerofmovement。LastnightIputonarowoflittlefliesnearoneedgeoftwoYOUNGISHleaves;andafter14hourstheseedgesarebeautifullyfoldedoversoastoclasptheflies,thusbringingtheglandsintocontactwiththeuppersurfacesoftheflies,andtheyarenowsecretingcopiouslyaboveandbelowthefliesandnodoubtabsorbing。Theacidsecretionhasrundownthechannellededgeandhascollectedinthespoon—shapedextremity,wherenodoubttheglandsareabsorbingthedelicioussoup。Theleafononesidelooksjustlikethehelixofahumanear,ifyouweretostuffflieswithinthefold。Yoursmostsincerely,CH。DARWIN。
  CHARLESDARWINTOASAGRAY。
  Down,June3[1874]……IamnowhardatworkgettingmybookonDrosera&Co。readyfortheprinters,butitwilltakesometime,forIamalwaysfindingoutnewpointstoobserve。IthinkyouwillbeinterestedbymyobservationsonthedigestiveprocessinDrosera;thesecretioncontainsanacidoftheaceticseries,andsomefermentcloselyanalogousto,butnotidenticalwith,pepsin;forIhavebeenmakingalongseriesofcomparativetrials。
  NohumanbeingwillbelievewhatIshallpublishaboutthesmallnessofthedosesofphosphateofammoniawhichact……IbeganreadingtheMadagascarsquib(Adescriptionofacarnivorousplantsupposedtosubsistonhumanbeings。)quitegravely,andwhenIfounditstatedthatFelisandBosinhabitedMadagascar,Ithoughtitwasafalsestory,anddidnotperceiveitwasahoaxtillIcametothewoman……
  CHARLESDARWINTOF。C。DONDERS。(ProfessorDonders,thewell—knownphysiologistofUtrecht。)
  Down,July7,1874。
  MydearProfessorDonders,MysonGeorgewritestomethathehasseenyou,andthatyouhavebeenverykindtohim,forwhichIreturntoyoumycordialthanks。Hetellsmeonyourauthority,ofafactwhichinterestsmeinthehighestdegree,andwhichImuchwishtobeallowedtoquote。Itrelatestotheactionofonemillionthofagrainofatropineontheeye。Nowwillyoubesokind,wheneveryoucanfindalittleleisure,totellmewhetheryouyourselfhaveobservedthisfact,orbelieveitongoodauthority。Ialsowishtoknowwhatproportionbyweighttheatropineboretothewatersolution,andhowmuchofthesolutionwasappliedtotheeye。ThereasonwhyIamsoanxiousonthisheadisthatitgivessomesupporttocertainfactsrepeatedlyobservedbymewithrespecttotheactionofphosphateofammoniaonDrosera。The1/4000000ofagrainabsorbedbyaglandclearlymakesthetentaclewhichbearsthisglandbecomeinflected;andIamfullyconvincedthat1/20000000ofagrainofthecrystallisedsalt(i。e。
  containingaboutone—thirdofitsweightofwaterofcrystallisation)doesthesame。NowIamquiteunhappyatthethoughtofhavingtopublishsuchastatement。Itwillbeofgreatvaluetometobeabletogiveanyanalogousfactsinsupport。ThecaseofDroseraisallthemoreinterestingastheabsorptionofthesaltoranyotherstimulantappliedtotheglandcausesittotransmitamotorinfluencetothebaseofthetentaclewhichbearsthegland。
  Prayforgivemefortroublingyou,anddonottroubleyourselftoanswerthisuntilyourhealthisfullyre—established。
  Praybelieveme,Yoursverysincerely,CHARLESDARWIN。
  [Duringthesummerof1874hewasatworkonthegenusUtricularia,andhewrote(July16th)toSirJ。D。Hookergivingsomeaccountoftheprogressofhiswork:——
  "IamrathergladyouhavenotbeenabletosendUtricularia,forthecommonspecieshasdrivenF。andmealmostmad。ThestructureisMOST
  complex。ThebladderscatchamultitudeofEntomostraca,andlarvaeofinsects。Themechanismforcaptureisexcellent。Butthereismuchthatwecannotunderstand。FromwhatIhaveseento—day,Istronglysuspectthatitisnecrophagous,i。e。thatitcannotdigest,butabsorbsdecayingmatter。"
  HewasindebtedtoLadyDorothyNevillforspecimensofthecuriousUtriculariamontana,whichisnotaquaticliketheEuropeanspecies,butgrowsamongthemossanddebrisonthebranchesoftrees。Tothisspeciesthefollowingletterrefers:]
  CHARLESDARWINTOLADYDOROTHYNEVILL。
  DownSeptember18[1874]。
  DearLadyDorothyNevill,Iamsomuchobligedtoyou。IwassoconvincedthatthebladderswerewiththeleavesthatIneverthoughtofremovingthemoss,andthiswasverystupidofme。Thegreatsolidbladder—likeswellingsalmostonthesurfacearewonderfulobjects,butarenotthetruebladders。TheseI
  foundontherootsnearthesurface,anddowntoadepthoftwoinchesinthesand。Theyareastransparentasglass,from1/20to1/100ofaninchinsize,andhollow。TheyhavealltheimportantpointsofstructureofthebladdersofthefloatingEnglishspecies,andIfeltconfidentIshouldfindcapturedprey。AndsoIhavetomydelightintwobladders,withclearproofthattheyhadabsorbedfoodfromthedecayingmass。ForUtriculariaisacarrion—feeder,andnotstrictlycarnivorouslikeDrosera。
  Thegreatsolidbladder—likebodies,Ibelieve,arereservoirsofwaterlikeacamel’sstomach。AssoonasIhavemadeafewmoreobservations,I
  meantobesocruelastogiveyourplantnowater,andobservewhetherthegreatbladdersshrinkandcontainairinsteadofwater;Ishallthenalsowashallearthfromallroots,andseewhethertherearetruebladdersforcapturingsubterraneaninsectsdowntotheverybottomofthepot。Nowshallyouthinkmeverygreedy,ifIsaythatsupposingthespeciesisnotveryprecious,andyouhaveseveral,willyougivemeonemoreplant,andifso,pleasetosenditto"OrpingtonStation,S。E。R。,tobeforwardedbyfootmessenger。"
  IhavehardlyeverenjoyedadaymoreinmylifethanIhavethisday’swork;andthisIowetoyourLadyship’sgreatkindness。
  Theseedsareverycuriousmonsters;IfancyofsomeplantalliedtoMedicago,butIwillshowthemtoDr。Hooker。
  Yourladyship’sverygratefully,CH。DARWIN。
  CHARLESDARWINTOJ。D。HOOKER。
  Down,September30,1874。
  MydearH。,YourmagnificentpresentofAldrovandahasarrivedquitesafe。Ihaveenjoyedgreatlyagoodlookattheshutleaves,oneofwhichIcutopen。
  ItisanaquaticDionaea,whichhasacquiredsomestructuresidenticalwiththoseofUtricularia!
  IftheleavesopenandIcantransferthemopenunderthemicroscope,I
  willtrysomeexperiments,formortalmancannotresistthetemptation。IfIcannottransfer,Iwilldonothing,forotherwiseitwouldrequirehundredsofleaves。
  Youareagoodmantogivemesuchpleasure。
  Yoursaffectionately,C。DARWIN。
  [Themanuscriptof’InsectivorousPlants’wasfinishedinMarch1875。Heseemstohavebeenmorethanusuallyoppressedbythewritingofthisbook,thushewrotetoSirJ。D。HookerinFebruary:——
  "Youaskaboutmybook,andallthatIcansayisthatIamreadytocommitsuicide;Ithoughtitwasdecentlywritten,butfindsomuchwantsrewriting,thatitwillnotbereadytogotoprintersfortwomonths,andwillthenmakeaconfoundedlybigbook。Murraywillsaythatitisnousepublishinginthemiddleofsummer,soIdonotknowwhatwillbetheupshot;butIbegintothinkthateveryonewhopublishesabookisafool。"
  ThebookwaspublishedonJuly2nd,1875,and2700copiesweresoldoutoftheeditionof3000。]
  CHAPTER2。XIV。
  THE’POWEROFMOVEMENTINPLANTS。’
  1880。
  [Thefewsentencesintheautobiographicalchaptergivewithsufficientclearnesstheconnectionbetweenthe’PowerofMovement,’andoneoftheauthor’searlierbooks,thaton’ClimbingPlants。’Thecentralideaofthebookisthatthemovementsofplantsinrelationtolight,gravitation,etc。,aremodificationsofaspontaneoustendencytorevolveorcircumnutate,whichiswidelyinherentinthegrowingpartsofplants。
  Thisconceptionhasnotbeengenerallyadopted,andhasnottakenaplaceamongthecanonsoforthodoxphysiology。ThebookhasbeentreatedbyProfessorSachswithafewwordsofprofessorialcontempt;andbyProfessorWiesnerithasbeenhonouredbycarefulandgenerouslyexpressedcriticism。
  Mr。ThiseltonDyer(’CharlesDarwin’(’Nature’Series),page41。)haswellsaid:"Whetherthismasterlyconceptionoftheunityofwhathashithertoseemedachaosofunrelatedphenomenawillbesustained,timealonewillshow。ButnoonecandoubttheimportanceofwhatMr。Darwinhasdone,inshowingthatforthefuturethephenomenaofplantmovementcanandindeedmustbestudiedfromasinglepointofview。"
  Theworkwasbeguninthesummerof1877,afterthepublicationof’DifferentFormsofFlowers,’andbytheautumnhisenthusiasmforthesubjectwasthoroughlyestablished,andhewrotetoMr。Dyer:"Iamallonfireatthework。"Atthistimehewasstudyingthemovementsofcotyledons,inwhichthesleepofplantsistobeobservedinitssimplestform;inthefollowingspringhewastryingtodiscoverwhatusefulpurposethesesleep—movementscouldserve,andwrotetoSirJosephHooker(March25th,1878):——
  "IthinkwehavePROVEDthatthesleepofplantsistolessentheinjurytotheleavesfromradiation。Thishasinterestedmemuch,andhascostusgreatlabour,asithasbeenaproblemsincethetimeofLinnaeus。Butwehavekilledorbadlyinjuredamultitudeofplants:N。B。——Oxaliscarnosawasmostvaluable,butlastnightwaskilled。"
  Hislettersofthisperioddonotgiveanyconnectedaccountoftheprogressofthework。Thetwofollowingaregivenasbeingcharacteristicoftheauthor:]
  CHARLESDARWINTOW。THISELTONDYER。
  Down,June2,1878。
  MydearDyer,IremembersayingthatIshoulddieadisgracedmanifIdidnotobserveaseedlingCactusandCycas,andyouhavesavedmefromthishorriblefate,astheymovesplendidlyandnormally。ButIhavetwoquestionstoask:
  theCycasobservedwasahugeseedinabroadandveryshallowpotwithcocoa—nutfibreasIsuppose。ItwasnamedonlyCycas。WasitCycaspectinata?IsupposethatIcannotbewronginbelievingthatwhatfirstappearsabovegroundisatrueleaf,forIcanseenostemoraxis。
  Lastly,youmayrememberthatIsaidthatwecouldnotraiseOpuntianigricans;nowImustconfesstoapieceofstupidity;onedidcomeup,butmygardenerandselfstaredatit,andconcludedthatitcouldnotbeaseedlingOpuntia,butnowthatIhaveseenoneofO。basilaris,Iamsureitwas;Iobserveditonlycasually,andsawmovements,whichmakesmewishtoobservecarefullyanother。Ifyouhaveanyfruit,willMr。Lynch(Mr。
  R。I。Lynch,nowCuratoroftheBotanicGardenatCambridgewasatthistimeintheRoyalGardens,Kew。)besokindastosendonemore?
  Iamworkingawaylikeaslaveatradicles[roots]andatmovementsoftrueleaves,forIhaveprettywelldonewithcotyledons……
  ThatwasanEXCELLENTletterabouttheGardens(ThisreferstoanattempttoinducetheGovernmenttoopentheRoyalGardensatKewinthemorning。):
  Ihadhopedthattheagitationwasover。Politiciansareapoortrucklinglot,for[they]mustseethewretchedeffectsofkeepingthegardensopenalldaylong。
  Yourevertroublesomefriend,CH。DARWIN。
  CHARLESDARWINTOW。THISELTONDYER。
  4BryanstonSt。,PortmanSquare,November21[1878]。
  MydearDyer,ImustthankyouforallthewonderfultroublewhichyouhavetakenabouttheseedsofImpatiens,andonscoresofotheroccasions。Itintruthmakesmefeelashamedofmyself,andIcannothelpthinking:"OhLord,whenheseesourbookhewillcryout,isthisallforwhichIhavehelpedsomuch!"Inseriousness,Ihopethatwehavemadeoutsomepoints,butI
  fearthatwehavedoneverylittleforthelabourwhichwehaveexpendedonourwork。Wearehereforaweekforalittlerest,whichIneeded。
  IfIrememberright,November30th,istheanniversaryattheRoyal,andI
  fearSirJosephmustbealmostatthelastgasp。IshallbegladwhenheisnolongerPresident。
  Yoursverysincerely,CH。DARWIN。
  [Inthespringofthefollowingyear,1879。Whenhewasengagedinputtinghisresultstogether,hewrotesomewhatdespondinglytoMr。Dyer:"Iamoverwhelmedwithmynotes,andalmosttoooldtoundertakethejobwhichI
  haveinhand——i。e。movementsofallkinds。Yetitisworsetobeidle。"
  Lateronintheyear,whentheworkwasapproachingcompletion,hewrotetoProf。Carus(July17,1879),withrespecttoatranslation:——
  "TogetherwithmysonFrancis,IampreparingaratherlargevolumeonthegeneralmovementsofPlants,andIthinkthatwehavemadeoutagoodmanynewpointsandviews。
  "IfearthatourviewswillmeetagooddealofoppositioninGermany;butwehavebeenworkingveryhardforsomeyearsatthesubject。
  "IshallbeMUCHpleasedifyouthinkthebookworthtranslating,andproof—sheetsshallbesentyou,whenevertheyareready。"
  Intheautumnhewashardatworkonthemanuscript,andwrotetoDr。Gray(October24,1879):——
  "Ihavewrittenaratherbigbook——moreisthepity——onthemovementsofplants,andIamnowjustbeginningtogoovertheMS。forthesecondtime,whichisahorridbore。"
  Onlytheconcludingpartofthenextletterreferstothe’PowerofMovements’:]
  CHARLESDARWINTOA。DECANDOLLE。
  May28,1880。
  MydearSir,Iamparticularlyobligedtoyouforhavingsokindlysendmeyour’Phytographie’(Abookonthemethodsofbotanicalresearch,moreespeciallyofsystematicwork。);forifIhadmerelyseenitadvertised,I
  shouldnothavesupposedthatitcouldhaveconcernedme。Asitis,Ihavereadwithverygreatinterestaboutaquarter,butwillnotdelaylongerthankingyou。Allthatyousayseemstomeveryclearandconvincing,andasinallyourwritingsIfindalargenumberofphilosophicalremarksnewtome,andnodoubtshallfindmanymore。TheyhaverecalledmanyapuzzlethroughwhichIpassedwhenmonographingtheCirripedia;andyourbookinthosedayswouldhavebeenquiteinvaluabletome。IthaspleasedmetofindthatIhavealwaysfollowedyourplanofmakingnotesonseparatepiecesofpaper;Ikeepseveralscoresoflargeportfolios,arrangedonverythinshelvesabouttwoinchesapart,fastenedtothewallsofmystudy,andeachshelfhasitspropernameortitle;andIcanthusputatonceeverymemorandumintoitsproperplace。Yourbookwill,Iamsure,beveryusefultomanyyoungstudents,andIshallbegmysonFrancis(whointendstodevotehimselftothephysiologyofplants)toreaditcarefully。