ThecaseofNewZealand,towhichfineislandweasyetowenowidelycultivatedplant,mayseemopposedtothisview;for,whenfirstdiscovered,thenativescultivatedseveralplants;butallinquirersbelieve,inaccordancewiththetraditionsofthenatives,thattheearlyPolynesiancolonistsbroughtwiththemseedsandroots,aswellasthedog,whichhadbeenwiselypreservedduringtheirlongvoyage。ThePolynesiansaresofrequentlylostontheoceanthatthisdegreeofprudencewouldoccurtoanywanderingparty:hencetheearlycolonistsofNewZealand,likethelaterEuropeancolonists,wouldnothavehadanystronginducementtocultivatetheaboriginalplants。AccordingtoDeCandolleweowethirty—
  threeusefulplantstoMexico,Peru,andChile;noristhissurprisingwhenwerememberthecivilisedstateoftheinhabitants,asshownbythefactoftheirhavingpractisedartificialirrigationandmadetunnelsthroughhardrockswithouttheuseofironorgunpowder,andwho,asweshallseeinafuturechapter,fullyrecognised,asfarasanimalswereconcerned,andthereforeprobablyinthecaseofplants,theimportantprincipleofselection。WeowesomeplantstoBrazil;andtheearlyvoyagers,namely,VespuciusandCabral,describethecountryasthicklypeopledandcultivated。InNorthAmerica(9/16。ForCanadaseeJ。Cartier’sVoyagein1534;forFloridaseeNarvaezandFerdinanddeSoto’sVoyages。AsIhaveconsultedtheseandotheroldVoyagesinmorethanonegeneralcollectionofVoyages,Idonotgiveprecisereferencestothepages。SeealsoforseveralreferencesAsaGrayinthe’AmericanJournalofScience’volume24
  November1857page441。ForthetraditionsofthenativesofNewZealandseeCrawfurd’GrammarandDict。oftheMalayLanguage’1852page260。)thenativescultivatedmaize,pumpkins,gourds,beans,andpeas,"alldifferentfromours,"andtobacco;andwearehardlyjustifiedinassumingthatnoneofourpresentplantsaredescendedfromtheseNorthAmericanforms。HadNorthAmericabeencivilisedforaslongaperiod,andasthicklypeopled,asAsiaorEurope,itisprobablethatthenativevines,walnuts,mulberries,crabs,andplums,wouldhavegivenrise,afteralongcourseofcultivation,toamultitudeofvarieties,someextremelydifferentfromtheirparent—stocks;andescapedseedlingswouldhavecausedintheNew,asintheOldWorld,muchperplexitywithrespecttotheirspecificdistinctnessandparentage。’(9/17。SeeforexampleMr。HewettC。Watson’sremarksonourwildplumsandcherriesandcrabs:’CybeleBritannica’
  volume1pages330,334,etc。VanMons(inhis’ArbresFruitiers’1835tome1page444)declaresthathehasfoundthetypesofallourcultivatedvarietiesinwildseedlings,butthenhelooksontheseseedlingsassomanyaboriginalstocks。)
  [CEREALIA。
  Iwillnowenterondetails。ThecerealscultivatedinEuropeconsistoffourgenera——wheat,rye,barley,andoats。Ofwheatthebestmodernauthorities(9/18。SeeA。DeCandolle’Geograph。Bot。’1855page928etseq。Godron’Del’Espece’1859tome2page70;andMetzger’DieGetreidearten’etc。1841。)makefourorfive,orevensevendistinctspecies;ofrye,one;ofbarley,three;andofoats,two,three,orfourspecies。Sothataltogetherourcerealsarerankedbydifferentauthorsunderfromtentofifteendistinctspecies。Thesehavegivenrisetoamultitudeofvarieties。Itisaremarkablefactthatbotanistsarenotuniversallyagreedontheaboriginalparent—formofanyonecerealplant。
  Forinstance,ahighauthoritywritesin1855(9/19。Mr。Benthaminhisreviewentitled’Hist。NotesoncultivatedPlants’byDr。A。Targioni—
  Tozzettiin’JournalofHort。Soc。’volume91855page133。Heinformsmethathestillretainsthesameopinion。),"Weourselveshavenohesitationinstatingourconviction,astheresultofallthemostreliableevidence,thatnoneoftheseCerealiaexist,orhaveexisted,trulywildintheirpresentstate,butthatallarecultivatedvarietiesofspeciesnowgrowingingreatabundanceinS。EuropeorW。Asia。"Ontheotherhand,Alph。DeCandolle(9/20。’Geograph。Bot。’page928。Thewholesubjectisdiscussedwithadmirablefulnessandknowledge。)hasadducedabundantevidencethatcommonwheat(Triticumvulgare)hasbeenfoundwildinvariouspartsofAsia,whereitisnotlikelytohaveescapedfromcultivation:andthereissomeforceinM。Godron’sremark,that,supposingtheseplantstobeescapedseedlings(9/21。Godron’Del’Espece’tome2page72。Afewyearsagotheexcellent,thoughmisinterpreted,observationsofM。FabreledmanypersonstobelievethatwheatwasamodifieddescendantofAegilops;butM。
  Godron(tome1page165)hasshownbycarefulexperimentsthatthefirststepintheseries,viz。Aegilopstriticoides,isahybridbetweenwheatandAe。ovata。Thefrequencywithwhichthesehybridsspontaneouslyarise,andthegradualmannerinwhichtheAe。triticoidesbecomesconvertedintotruewheat,aloneleaveanydoubtwithrespecttoM。Godron’sconclusions。),astheyhavepropagatedthemselvesinawildstateforseveralgenerations,theircontinuedresemblancetocultivatedwheatrendersitprobablethatthelatterhasretaineditsaboriginalcharacter。
  Butthestrongtendencytoinheritance,whichmostofthevarietiesofwheatevince,asweshallpresentlysee,isheregreatlyundervalued。MuchweightmustalsobeattributedtoaremarkbyProfessorHildebrand(9/22。
  ’DieVerbreitungsmittelderPflanzen’1873page129。)thatwhentheseedsorfruitofcultivatedplantspossessqualitiesdisadvantageoustothemasameansofdistribution,wemayfeelalmostsurethattheynolongerretaintheiraboriginalcondition。Ontheotherhand,M。DeCandolleinsistsstronglyonthefrequentoccurrenceintheAustriandominionsofryeandofonekindofoatsinanapparentlywildcondition。Withtheexceptionofthesetwocases,whichhoweverareratherdoubtful,andwiththeexceptionoftwoformsofwheatandoneofbarley,whichhebelievestohavebeenfoundtrulywild,M。DeCandolledoesnotseemfullysatisfiedwiththeotherreporteddiscoveriesoftheparent—formsofourothercereals。Withrespecttooats,accordingtoMr。Buckmann(9/23。ReporttoBritishAssociationfor1857page207。),thewildEnglishAvenafatuacanbeconvertedbyafewyearsofcarefulcultivationandselectionintoformsalmostidenticalwithtwoverydistinctcultivatedraces。Thewholesubjectoftheoriginandspecificdistinctnessofthevariouscerealplantsisamostdifficultone;butweshallperhapsbeabletojudgealittlebetterafterconsideringtheamountofvariationwhichwheathasundergone。
  Metzgerdescribessevenspeciesofwheat,Godronreferstofive,andDeCandolletoonlyfour。Itisnotimprobablethat,besidesthekindsknowninEurope,otherstronglycharacterisedformsexistinthemoredistantpartsoftheworld;forLoiseleur—Deslongchamps(9/24。ConsiderationssurlesCereales’1842—43page29。)speaksofthreenewspeciesorvarieties,senttoEuropein1822fromChineseMongolia,whichheconsidersasbeingthereindigenous。Moorcroft(9/25。’TravelsintheHimalayanProvinces’
  etc。1841volume1page224。)alsospeaksofHasorawheatinLadakhasverypeculiar。Ifthosebotanistsarerightwhobelievethatatleastsevenspeciesofwheatoriginallyexisted,thentheamountofvariationinanyimportantcharacterwhichwheathasundergoneundercultivationhasbeenslight;butifonlyfouroralessernumberofspeciesoriginallyexisted,thenitisevidentthatvarietieshavearisensostronglymarked,thattheyhavebeenconsideredbycapablejudgesasspecificallydistinct。Buttheimpossibilityofdecidingwhichformsoughttoberankedasspeciesandwhichasvarieties,makesituselesstospecifyindetailthedifferencesbetweenthevariouskindsofwheat。Speakinggenerally,theorgansofvegetationdifferlittle(9/26。Col。J。LeCouteuronthe’VarietiesofWheat’pages23,79。);butsomekindsgrowcloseandupright,whilstothersspreadandtrailalongtheground。Thestrawdiffersinbeingmoreorlesshollow,andinquality。Theears(9/27。Loiseleur—Deslongchamps’Consid。
  surlesCereales’page11。)differincolourandinshape,beingquadrangular,compressed,ornearlycylindrical;andthefloretsdifferintheirapproximationtoeachother,intheirpubescence,andinbeingmoreorlesselongated。Thepresenceorabsenceofbarbsisaconspicuousdifference,andincertainGramineaeservesevenasagenericcharacter(9/28。SeeanexcellentreviewinHooker’Journ。ofBotany’volume8page82note。);although,asremarkedbyGodron(9/29。’Del’Espece’tome2page73。)thepresenceofbarbsisvariableincertainwildgrasses,andespeciallyinthosesuchasBromussecalinusandLoliumtemulentum,whichhabituallygrowmingledwithourcerealcrops,andwhichhavethusunintentionallybeenexposedtoculture。Thegrainsdifferinsize,weight,andcolour;inbeingmoreorlessdownyatoneend,inbeingsmoothorwrinkled,inbeingeithernearlyglobular,oval,orelongated;andfinallyininternaltexture,beingtenderorhard,orevenalmosthorny,andintheproportionofglutenwhichtheycontain。
  Nearlyalltheracesorspeciesofwheatvary,asGodron(9/30。Ibidtome2
  page75。)hasremarked,inanexactlyparallelmanner,——intheseedbeingdownyorglabrous,andincolour,——andinthefloretsbeingbarbedornotbarbed,etc。Thosewhobelievethatallthekindsaredescendedfromasinglewildspeciesmayaccountforthisparallelvariationbytheinheritanceofasimilarconstitution,andaconsequenttendencytovaryinthesamemanner;andthosewhobelieveinthegeneraltheoryofdescentwithmodificationmayextendthisviewtotheseveralspeciesofwheat,ifsucheverexistedinastateofnature。
  Althoughfewofthevarietiesofwheatpresentanyconspicuousdifference,theirnumberisgreat。Dalbretcultivatedduringthirtyyearsfrom150to160kinds,andexceptinginthequalityofthegraintheyallkepttrue;
  ColonelLeCouteurpossessedupwardsof150,andPhilippar322varieties。
  (9/31。ForDalbretandPhilipparseeLoiseleur—Deslongchamps’Consid。surlesCereales’pages45,70。LeCouteuronWheatpages6,14—17。)Aswheatisanannual,wethusseehowstrictlymanytriflingdifferencesincharacterareinheritedthroughmanygenerations。ColonelLeCouteurinsistsstronglyonthissamefact。Inhisperseveringandsuccessfulattemptstoraisenewvarieties,hefoundthattherewasonlyone"securemodetoensurethegrowthofpuresorts,namely,togrowthemfromsinglegrainsorfromsingleears,andtofollowuptheplanbyafterwardssowingonlytheproduceofthemostproductivesoastoformastock。"ButMajorHallett(9/32。SeehisEssayon’PedigreeinWheat’1862;alsopaperreadbeforetheBritishAssociation1869andotherpublications。)hasgonemuchfarther,andbythecontinuedselectionofplantsfromthegrainsofthesameear,duringsuccessivegenerations,hasmadehis’PedigreeinWheat’
  (andothercereals)nowfamousinmanyquartersoftheworld。Thegreatamountofvariabilityintheplantsofthesamevarietyisanotherinterestingpoint,whichwouldneverhavebeendetectedexceptbyaneyelongpractisedtothework;thusColonelLeCouteurrelates(9/33。
  ’VarietiesofWheat’Introductionpage6。Marshallinhis’RuralEconomyofYorkshire’volume2page9remarksthat"ineveryfieldofcornthereisasmuchvarietyasinaherdofcattle。")thatinafieldofhisownwheat,whichheconsideredatleastaspureasthatofanyofhisneighbours,ProfessorLaGascafoundtwenty—threesorts;andProfessorHenslowhasobservedsimilarfacts。Besidessuchindividualvariations,formssufficientlywellmarkedtobevaluedandtobecomewidelycultivatedsometimessuddenlyappear:thusMr。Shirreffhashadthegoodfortunetoraiseinhislifetimesevennewvarieties,whicharenowextensivelygrowninmanypartsofBritain。(9/34。’Gardener’sChronicle’and’Agricult。
  Gazette’1862page963。)
  Asinthecaseofmanyotherplants,somevarieties,botholdandnew,arefarmoreconstantincharacterthanothers。ColonelLeCouteurwasforcedtorejectsomeofhisnewsub—varieties,whichhesuspectedhadbeenproducedfromacross,asincorrigiblysportive。OntheotherhandMajorHallett(9/35。’Gardener’sChronicle’November1868page1199。)hasshownhowwonderfullyconstantsomevarietiesare,althoughnotancientones,andalthoughcultivatedinvariouscountries。Withrespecttothetendencytovary,Metzger(9/36。’Getreidearten’1841s。66,91,92,116,117。)givesfromhisownexperiencesomeinterestingfacts:hedescribesthreeSpanishsub—varieties,moreespeciallyoneknowntobeconstantinSpain,whichinGermanyassumedtheirpropercharacteronlyduringhotsummers;anothervarietykepttrueonlyingoodland,butafterhavingbeencultivatedfortwenty—fiveyearsbecamemoreconstant。Hementionstwoothersub—varietieswhichwereatfirstinconstant,butsubsequentlybecame,apparentlywithoutanyselection,accustomedtotheirnewhomes,andretainedtheirpropercharacter。Thesefactsshowwhatsmallchangesintheconditionsoflifecausevariability,andtheyfurthershowthatavarietymaybecomehabituatedtonewconditions。OneisatfirstinclinedtoconcludewithLoiseleur—Deslongchamps,thatwheatcultivatedinthesamecountryisexposedtoremarkablyuniformconditions;butmanuresdiffer,seedistakenfromonesoiltoanother,and,whatisfarmoreimportant,theplantsareexposedaslittleaspossibletostrugglewithotherplants,andarethusenabledtoexistunderdiversifiedconditions。Inastateofnatureeachplantisconfinedtothatparticularstationandkindofnutrimentwhichitcanseizefromtheotherplantsbywhichitissurrounded。
  Wheatquicklyassumesnewhabitsoflife。ThesummerandwinterkindswereclassedbyLinnaeusasdistinctspecies;butM。Monnier(9/37。QuotedbyGodron’Del’Espece’volume2page74。Soitis,accordingtoMetzger’Getreidearten’s。18,withsummerandwinterbarley。)hasprovedthatthedifferencebetweenthemisonlytemporary。Hesowedwinter—wheatinspring,andoutofonehundredplantsfouraloneproducedripeseeds;theseweresownandresown,andinthreeyearsplantswererearedwhichripenedalltheirseed。Conversely,nearlyalltheplantsraisedfromsummer—wheat,whichwassowninautumn,perishedfromfrost;butafewweresavedandproducedseed,andinthreeyearsthissummer—varietywasconvertedintoawinter—variety。Henceitisnotsurprisingthatwheatsoonbecomestoacertainextentacclimatised,andthatseedbroughtfromdistantcountriesandsowninEuropevegetatesatfirst,orevenforaconsiderableperiod(9/38。Loiseleur—Deslongchamps’Cereales’part2page224。LeCouteurpage70。Manyotheraccountscouldbeadded。)differentlyfromourEuropeanvarieties。InCanadathefirstsettlers,accordingtoKalm(9/39。’TravelsinNorthAmerica’1753—1761Englishtranslationvolume3page165。),foundtheirwinterstoosevereforwinter—wheatbroughtfromFrance,andtheirsummersoftentooshortforsummer—wheat;andtheythoughtthattheircountrywasuselessforcorncropsuntiltheyprocuredsummer—wheatfromthenorthernpartsofEurope,whichsucceededwell。Itisnotoriousthattheproportionofglutendiffersmuchunderdifferentclimates。Theweightofthegrainisalsoquicklyaffectedbyclimate:Loiseleur—Deslongchamps(9/40。’Cereales’part2pages179—183。)sowednearParis54varieties,obtainedfromtheSouthofFranceandfromtheBlackSea,and52oftheseyieldedseedfrom10to40percentheavierthantheparent—seed。HethensenttheseheaviergrainsbacktotheSouthofFrance,buttheretheyimmediatelyyieldedlighterseed。
  Allthosewhohavecloselyattendedtothesubjectinsistonthecloseadaptationofnumerousvarietiesofwheattovarioussoilsandclimatesevenwithinthesamecountry;thusColonelLeCouteur(9/41。’OntheVarietiesofWheat’Introductionpage7。SeeMarshall’RuralEcon。ofYorkshire’volume2page9。Withrespecttosimilarcasesofadaptationinthevarietiesofoatsseesomeinterestingpapersinthe’Gardener’sChronicleandAgricult。Gazette’1850pages204,219。)says,"Itisthesuitablenessofeachsorttoeachsoilthatwillenablethefarmertopayhisrentbysowingonevariety,wherehewouldbeunabletodosobyattemptingtogrowanotherofaseeminglybettersort。"Thismaybeinpartduetoeachkindbecominghabituatedtoitsconditionsoflife,asMetzgerhasshowncertainlyoccurs,butitisprobablyinmainpartduetoinnatedifferencesbetweentheseveralvarieties。
  Muchhasbeenwrittenonthedeteriorationofwheat;thatthequalityoftheflour,sizeofgrain,timeofflowering,andhardness,maybemodifiedbyclimateandsoil,seemsnearlycertain;butthatthewholebodyofanyonesub—varietyeverbecomeschangedintoanotheranddistinctsub—variety,thereisnoreasontobelieve。Whatapparentlydoestakeplace,accordingtoLeCouteur(9/42。’OntheVarietiesofWheat’page59。Mr。Shirreffandahigherauthoritycannotbegiven(’Gardener’sChronicleandAgricult。
  Gazette’1862page963),says"Ihaveneverseengrainwhichhaseitherbeenimprovedordegeneratedbycultivation,soastoconveythechangetothesucceedingcrop。),is,thatsomeonesub—varietyoutofthemanywhichmayalwaysbedetectedinthesamefieldismoreprolificthantheothers,andgraduallysupplantsthevarietywhichwasfirstsown。
  Withrespecttothenaturalcrossingofdistinctvarietiestheevidenceisconflicting,butpreponderatesagainstitsfrequentoccurrence。Manyauthorsmaintainthatimpregnationtakesplaceintheclosedflower,butI
  amsurefrommyownobservationthatthisisnotthecase,atleastwiththosevarietiestowhichIhaveattended。ButasIshallhavetodiscussthissubjectinanotherwork,itmaybeherepassedover。]
  Inconclusion,allauthorsadmitthatnumerousvarietiesofwheathavearisen;buttheirdifferencesareunimportant,unless,indeed,someoftheso—calledspeciesarerankedasvarieties。ThosewhobelievethatfromfourtosevenwildspeciesofTriticumoriginallyexistedinnearlythesameconditionasatpresent,resttheirbeliefchieflyonthegreatantiquityoftheseveralforms。(9/43。Alph。DeCandolle’Geograph。Bot。’page930。)
  Itisanimportantfact,whichwehaverecentlylearntfromtheadmirableresearchesofHeer(9/44。’PflanzenderPfahlbauten’1866。),thattheinhabitantsofSwitzerland,evensoearlyastheNeolithicperiod,cultivatednolessthantencerealplants,namely,fivekindsofwheat,ofwhichatleastfourarecommonlylookedatasdistinctspecies,threekindsofbarley,apanicum,andasetaria。Ifitcouldbeshownthatattheearliestdawnofagriculturefivekindsofwheatandthreeofbarleyhadbeencultivated,weshouldofcoursebecompelledtolookattheseformsasdistinctspecies。But,asHeerhasremarked,agricultureevenattheNeolithicperiod,hadalreadymadeconsiderableprogress;for,besidesthecereals,peas,poppies,flax,andapparentlyapples,werecultivated。Itmayalsobeinferred,fromonevarietyofwheatbeingthesocalledEgyptian,andfromwhatisknownofthenativecountryofthepanicumandsetaria,aswellasfromthenatureoftheweedswhichthengrewmingledwiththecrops,thatthelake—inhabitantseitherstillkeptupcommercialintercoursewithsomesouthernpeopleorhadoriginallyproceededascolonistsfromtheSouth。
  Loiseleur—Deslongchamps(9/45。’LesCereales’page94。)hasarguedthat,ifourcerealplantshavebeengreatlymodifiedbycultivation,theweedswhichhabituallygrowmingledwiththemwouldhavebeenequallymodified。
  Butthisargumentshowshowcompletelytheprincipleofselectionhasbeenoverlooked。Thatsuchweedshavenotvaried,oratleastdonotvarynowinanyextremedegree,istheopinionofMr。H。C。WatsonandProfessorAsaGray,astheyinformme;butwhowillpretendtosaythattheydonotvaryasmuchastheindividualplantsofthesamesub—varietyofwheat?Wehavealreadyseenthatpurevarietiesofwheat,cultivatedinthesamefield,offermanyslightvariations,whichcanbeselectedandseparatelypropagated;andthatoccasionallymorestronglypronouncedvariationsappear,which,asMr。Shirreffhasproved,arewellworthyofextensivecultivation。Notuntilequalattentionbepaidtothevariabilityandselectionofweeds,cantheargumentfromtheirconstancyunderunintentionalculturebeofanyvalue。Inaccordancewiththeprinciplesofselectionwecanunderstandhowitisthatintheseveralcultivatedvarietiesofwheattheorgansofvegetationdiffersolittle;forifaplantwithpeculiarleavesappeared,itwouldbeneglectedunlessthegrainsofcornwereatthesametimesuperiorinqualityorsize。theselectionofseed—cornwasstronglyrecommended(9/46。QuotedbyLeCouteurpage16。)inancienttimesbyColumellaandCelsus;andasVirgilsays,——
  I’veseenthelargestseeds,tho’view’dwithcare,Degenerate,unlessth’industrioushandDidyearlycullthelargest。"
  Butwhetherinancienttimesselectionwasmethodicallypursuedwemaywelldoubt,whenwehearhowlaborioustheworkhasbeenfoundbyLeCoutourandHallett。Althoughtheprincipleofselectionissoimportant,yetthelittlewhichmanhaseffected,byincessantefforts(9/47。A。DeCandolle’Geograph。Bot。’page932。)duringthousandsofyears,inrenderingtheplantsmoreproductiveorthegrainsmorenutritiousthantheywereinthetimeoftheoldEgyptians,wouldseemtospeakstronglyagainstitsefficacy。Butwemustnotforgetthatateachsuccessiveperiodthestateofagricultureandthequantityofmanuresuppliedtothelandwillhavedeterminedthemaximumdegreeofproductiveness;foritwouldbeimpossibletocultivateahighlyproductivevariety,unlessthelandcontainedasufficientsupplyofthenecessarychemicalelements。
  Wenowknowthatmanwassufficientlycivilisedtocultivatethegroundatanimmenselyremoteperiod;sothatwheatmighthavebeenimprovedlongagouptothatstandardofexcellencewhichwaspossibleunderthethenexistingstateofagriculture。Onesmallclassoffactssupportsthisviewoftheslowandgradualimprovementofourcereals。Inthemostancientlake—habitationsofSwitzerland,whenmenemployedonlyflint—tools,themostextensivelycultivatedwheatwasapeculiarkind,withremarkablysmallearsandgrains。(9/48。O。Heer’DiePflanzenderPfahlbauten’1866。
  ThefollowingpassageisquotedfromDr。Christin’DieFaunaderPfahlbauten,vonDr。Rutimeyer’1861s。225。)"Whilstthegrainsofthemodernformsareinsectionfromseventoeightmillimetresinlength,thelargergrainsfromthelake—habitationsaresix,seldomseven,andthesmalleronesonlyfour。Theearisthusmuchnarrower,andthespikeletsstandoutmorehorizontally,thaninourpresentforms。"Soagainwithbarley,themostancientandmostextensivelycultivatedkindhadsmallears,andthegrainswere"smaller,shorter,andnearertoeachother,thaninthatnowgrown;withoutthehusktheywere21/2lineslong,andscarcely11/2broad,whilstthosenowgrownhavealengthofthreelines,andalmostthesameinbreadth。"(9/49。HeerasquotedbyCarlVogt’LecturesonMan’Englishtranslationpage355。)Thesesmall—grainedvarietiesofwheatandbarleyarebelievedbyHeertobetheparent—formsofcertainexistingalliedvarieties,whichhavesupplantedtheirearlyprogenitors。
  HeergivesaninterestingaccountofthefirstappearanceandfinaldisappearanceoftheseveralplantswhichwerecultivatedingreaterorlessabundanceinSwitzerlandduringformersuccessiveperiods,andwhichgenerallydifferedmoreorlessfromourexistingvarieties。Thepeculiarsmall—earedandsmall—grainedwheat,alreadyalludedto,wasthecommonestkindduringtheStoneperiod;itlasteddowntotheHelvetico—Romanage,andthenbecameextinct。Asecondkindwasrareatfirst,butafterwardsbecamemorefrequent。Athird,theEgyptianwheat(T。turgidum),doesnotagreeexactlywithanyexistingvariety,andwasrareduringtheStoneperiod。Afourthkind(T。dicoccum)differsfromallknownvarietiesofthisform。Afifthkind(T。monococcum)isknowntohaveexistedduringtheStoneperiodonlybythepresenceofasingleear。Asixthkind,thecommonT。spelta,wasnotintroducedintoSwitzerlanduntiltheBronzeage。Ofbarley,besidestheshort—earedandsmall—grainedkind,twootherswerecultivated,oneofwhichwasveryscarce,andresembledourpresentcommonH。distichum。DuringtheBronzeageryeandoatswereintroduced;theoat—
  grainsbeingsomewhatsmallerthanthoseproducedbyourexistingvarieties。ThepoppywaslargelycultivatedduringtheStoneperiod,probablyforitsoil;butthevarietywhichthenexistedisnotnowknown。
  ApeculiarpeawithsmallseedslastedfromtheStonetotheBronzeage,andthenbecameextinct;whilstapeculiarbean,likewisehavingsmallseeds,cameinattheBronzeperiodandlastedtothetimeoftheRomans。
  Thesedetailssoundlikethedescriptionsgivenbypalaeontologistsofthefirstappearance,theincreasingrarity,andfinalextinctionormodificationoffossilspecies,embeddedinthesuccessivestagesofageologicalformation。
  Finally,everyonemustjudgeforhimselfwhetheritismoreprobablethattheseveralformsofwheat,barley,rye,andoatsaredescendedfrombetweentenandfifteenspecies,mostofwhicharenoweitherunknownorextinct,orwhethertheyaredescendedfrombetweenfourandeightspecies,whichmayhaveeithercloselyresembledourpresentcultivatedforms,orhavebeensowidelydifferentastoescapeidentification。Inthislattercasewemustconcludethatmancultivatedthecerealsatanenormouslyremoteperiod,andthatheformerlypractisedsomedegreeofselection,whichinitselfisnotimprobable。Wemay,perhaps,furtherbelievethat,whenwheatwasfirstcultivatedtheearsandgrainsincreasedquicklyinsize,inthesamemannerastherootsofthewildcarrotandparsnipareknowntoincreasequicklyinbulkundercultivation。
  [MAIZEORINDIANCORN:Zeamays。
  Botanistsarenearlyunanimousthatallthecultivatedkindsbelongtothesamespecies。Itisundoubtedly(9/50。SeeAlph。DeCandolle’slongdiscussioninhis’Geograph。Bot。’page942。WithrespecttoNewEnglandseeSilliman’s’AmericanJournal’volume44page99。)ofAmericanorigin,andwasgrownbytheaboriginesthroughoutthecontinentfromNewEnglandtoChili。Itscultivationmusthavebeenextremelyancient,forTschudi(9/51。’TravelsinPeru’Englishtranslationpage177。)describestwokinds,nowextinctornotknowninPeru,whichweretakenfromtombsapparentlypriortothedynastyoftheIncas。’Butthereisevenstrongerevidenceofantiquity,forIfoundonthecoastofPeru(9/52。’Geolog。
  Observ。onS。America’1846page49。)headsofmaize,togetherwitheighteenspeciesofrecentsea—shell,embeddedinabeachwhichhadbeenupraisedatleast85feetabovethelevelofthesea。Inaccordancewiththisancientcultivation,numerousAmericanvarietieshavearisen。Theaboriginalformhasnotasyetbeendiscoveredinthewildstate。A
  peculiarkind(9/53。ThismaizeisfiguredinBonafous’magnificentwork,’Hist。Nat。duMais’1836P1。v。bis,andinthe’JournalofHort。Soc。’
  volume11846page115whereanaccountisgivenoftheresultofsowingtheseed。AyoungGuaranyIndian,onseeingthiskindofmaize,toldAugusteSt。Hilaire(seeDeCandolle’Geograph。Bot。’page951)thatitgrewwildinthehumidforestsofhisnativeland。Mr。Teschemacherin’Proc。BostonSoc。Hist。’October19,1842givesanaccountofsowingtheseed。),inwhichthegrains,insteadofbeingnaked,areconcealedbyhusksasmuchaselevenlinesinlength,hasbeenstated,butoninsufficientevidence,togrowwildinBrazil。Itisalmostcertainthattheaboriginalformwouldhavehaditsgrainsthusprotected(9/54。Moquin—Tandon’ElementsdeTeratologie’1841page126。);buttheseedsoftheBrazilianvarietyproduce,asIhearfromProfessorAsaGray,andasisstatedintwopublishedaccounts,eithercommonorhuskedmaize;anditisnotcrediblethatawildspecies,whenfirstcultivated,shouldvarysoquicklyandinsogreatadegree。
  Maizehasvariedinanextraordinaryandconspicuousmanner。Metzger(9/55。
  ’DieGetreidearten’1841s。208。IhavemodifiedafewofMetzger’sstatementsinaccordancewiththosemadebyBonafousinhisgreatwork’Hist。Nat。duMais’1836。),whopaidparticularattentiontothecultivationofthisplant,makestwelveraces(unter—art)withnumeroussub—varieties:ofthelattersomearetolerablyconstant,othersquiteinconstant。Thedifferentracesvaryinheightfrom15—18feettoonly16—
  18inches,asinadwarfvarietydescribedbyBonafous。Thewholeearisvariableinshape,beinglongandnarrow,orshortandthick,orbranched。
  Theearinonevarietyismorethanfourtimesaslongasinadwarfkind。
  Theseedsarearrangedintheearinfromsixtoeventwentyrows,orareplacedirregularly。Theseedsarecoloured——white,pale—yellow,orange,red,violet,orelegantlystreakedwithblack(9/56。Godron’Del’Espece’
  tome2page80;Al。DeCandolleibidpage951。);andinthesameeartherearesometimesseedsoftwocolours。InasmallcollectionIfoundthatasinglegrainofonevarietynearlyequalledinweightsevengrainsofanothervariety。Theshapeoftheseedvariesgreatly,beingveryflat,ornearlyglobular,oroval;broaderthanlong,orlongerthanbroad;withoutanypoint,orproducedintoasharptooth,andthistoothissometimesrecurved。Onevariety(therugosaofBonafous,andwhichisextensivelycultivatedintheUnitedStatesassweetcorn)hasitsseedscuriouslywrinkled,givingtothewholeearasingularappearance。Anothervariety(thecymosaofBon。)carriesitsearssocrowdedtogetherthatitiscalledmaisabouquet。Theseedsofsomevarietiescontainmuchglucoseinsteadofstarch。Maleflowerssometimesappearamongstthefemaleflowers,andMr。
  J。Scotthaslatelyobservedtherarercaseoffemaleflowersonatruemalepanicle,andlikewisehermaphroditeflowers。(9/57。’Transact。Bot。
  Soc。ofEdinburgh’volume8page60。)Azaradescribes(9/58。’Voyagesdansl’AmeriqueMeridionale’tome1page147。)avarietyinParaguaythegrainsofwhichareverytender,andhestatesthatseveralvarietiesarefittedforbeingcookedinvariousways。Thevarietiesalsodiffergreatlyinprecocity,andhavedifferentpowersofresistingdrynessandtheactionofviolentwind。(9/59。Bonafous’Hist。Nat。duMais’page31。)Someoftheforegoingdifferenceswouldcertainlybeconsideredofspecificvaluewithplantsinastateofnature。
  LeComteRestatesthatthegrainsofallthevarietieswhichhecultivatedultimatelyassumedayellowcolour。ButBonafous(9/60。Ibidpage31。)
  foundthatmostofthosewhichhesowedfortenconsecutiveyearskepttruetotheirpropertints;andheaddsthatinthevalleysofthePyreneesandontheplainsofPiedmontawhitemaizehasbeencultivatedformorethanacentury,andhasundergonenochange。
  Thetallkindsgrowninsouthernlatitudes,andthereforeexposedtogreatheat,requirefromsixtosevenmonthstoripentheirseed;whereasthedwarfkinds,growninnorthernandcolderclimates,requireonlyfromthreetofourmonths。(9/61。Metzger’Getreidearten’s。206。)PeterKalm(9/62。
  ’DescriptionofMaize’byP。Kalm1752in’SwedishActs’volume4。IhaveconsultedanoldEnglishMS。translation。),whoparticularlyattendedtothisplant,says,thatintheUnitedStates,inproceedingfromsouthtonorth,theplantssteadilydiminishinbulk。Seedsbroughtfromlat。37deginVirginia,andsowninlat。43—44deginNewEngland,produceplantswhichwillnotripentheirseed,orripenthemwiththeutmostdifficulty。
  SoitiswithseedcarriedfromNewEnglandtolat。45—47deginCanada。Bytakinggreatcareatfirst,thesouthernkindsaftersomeyears’cultureripentheirseedperfectlyintheirnorthernhomes,sothatthisisananalogouscasewiththatoftheconversionofsummerintowinterwheat,andconversely。Whentallanddwarfmaizeareplantedtogether,thedwarfkindsareinfullflowerbeforetheothershaveproducedasingleflower;andinPennsylvaniatheyripentheirseedssixweeksearlierthanthetallmaize。
  MetzgeralsomentionsaEuropeanmaizewhichripensitsseedfourweeksearlierthananotherEuropeankind。Withthesefacts,soplainlyshowinginheritedacclimatisation,wemayreadilybelieveKalm,whostatesthatinNorthAmericamaizeandsomeotherplantshavegraduallybeencultivatedfurtherandfurthernorthward。Allwritersagreethattokeepthevarietiesofmaizepuretheymustbeplantedseparatelysothattheyshallnotcross。
  TheeffectsoftheclimateofEuropeontheAmericanvarietiesishighlyremarkable。MetzgerobtainedseedfromvariouspartsofAmerica,andcultivatedseveralkindsinGermany。Iwillgiveanabstractofthechangesobserved(9/63。’Getreidearten’s。208。)inonecase,namely,withatallkind(Breit—kornigermais,Zeaaltissima)broughtfromthewarmerpartsofAmerica。Duringthefirstyeartheplantsweretwelvefeethigh,andafewseedswereperfected;thelowerseedsintheearkepttruetotheirproperform,buttheupperseedsbecameslightlychanged。Inthesecondgenerationtheplantswerefromninetotenfeetinheight,andripenedtheirseedbetter;thedepressionontheoutersideoftheseedhadalmostdisappeared,andtheoriginalbeautifulwhitecolourhadbecomeduskier。
  Someoftheseedshadevenbecomeyellow,andintheirnowroundedformtheyapproachedcommonEuropeanmaize。InthethirdgenerationnearlyallresemblancetotheoriginalandverydistinctAmericanparent—formwaslost。InthesixthgenerationthismaizeperfectlyresembledaEuropeanvariety,describedasthesecondsub—varietyofthefifthrace。WhenMetzgerpublishedhisbook,thisvarietywasstillcultivatednearHeidelberg,andcouldbedistinguishedfromthecommonkindonlybyasomewhatmorevigorousgrowth。AnalogousresultswereobtainedbythecultivationofanotherAmericanrace,the"white—toothcorn,"inwhichthetoothnearlydisappearedeveninthesecondgeneration。Athirdrace,the"chicken—corn,"didnotundergosogreatachange,buttheseedsbecamelesspolishedandpellucid。Intheabovecasestheseedswerecarriedfromawarmtoacolderclimate。ButFritzMullerinformsmethatadwarfvarietywithsmallroundedseeds(papa—gaien—mais),introducedfromGermanyintoS。Brazil,producesplantsastall,withseedsasflat,asthoseofthekindcommonlycultivatedthere。]
  Thesefactsaffordthemostremarkableinstanceknowntomeofthedirectandpromptactionofclimateonaplant。Itmighthavebeenexpectedthatthetallnessofthestem,theperiodofvegetation,andtheripeningoftheseed,wouldhavebeenthusaffected;butitisamuchmoresurprisingfactthattheseedsshouldhaveundergonesorapidandgreatachange。As,however,flowers,withtheirproducttheseed,areformedbythemetamorphosisofthestemandleaves,anymodificationintheselatterorganswouldbeapttoextend,throughcorrelation,totheorgansoffructification。
  [CABBAGE(Brassicaoleracea)。
  Everyoneknowshowgreatlythevariouskindsofcabbagedifferinappearance。IntheIslandofJersey,fromtheeffectsofparticularcultureandofclimateastalkhasgrowntotheheightofsixteenfeet,and"haditsspringshootsatthetopoccupiedbyamagpie’snest:"thewoodystemsarenotunfrequentlyfromtentotwelvefeetinheight,andarethereusedasrafters(9/64。’CabbageTimber’’Gardener’sChronicle’1856page744,quotedfromHooker’JournalofBotany。’Awalking—stickmadefromacabbage—stalkisexhibitedintheMuseumatKew。)andaswalking—sticks。WearethusremindedthatincertaincountriesplantsbelongingtothegenerallyherbaceousorderoftheCruciferaearedevelopedintotrees。
  Everyonecanappreciatethedifferencebetweengreenorredcabbageswithgreatsingleheads;Brussel—sproutswithnumerouslittleheads;broccolisandcauliflowerswiththegreaternumberoftheirflowersinanabortedcondition,incapableofproducingseed,andborneinadensecorymbinsteadofanopenpanicle;savoyswiththeirblisteredandwrinkledleaves;andborecolesandkails,whichcomenearesttothewildparent—form。Therearealsovariousfrizzledandlaciniatedkinds,someofsuchbeautifulcoloursthatVilmorininhisCatalogueof1851enumeratestenvarietieswhicharevaluedsolelyforornament。Somekindsarelesscommonlyknown,suchasthePortugueseCouveTronchuda,withtheribsofitsleavesgreatlythickened;
  andtheKohlrabiorchoux—raves,withtheirstemsenlargedintogreatturnip—likemassesabovetheground;andtherecentlyformednewrace(9/65。’JournaldelaSoc。Imp。d’Horticulture’1855page254quotedfrom’Gartenflora’April1855。)ofthechoux—raves,alreadyincludingninesub—
  varieties,inwhichtheenlargedpartliesbeneaththegroundlikeaturnip。
  Althoughweseesuchgreatdifferencesintheshape,size,colour,arrangement,andmannerofgrowthoftheleavesandstem,andoftheflower—stemsinthebroccoliandcauliflower,itisremarkablethattheflowersthemselves,theseed—podsandseeds,presentextremelyslightdifferencesornoneatall。(9/66。Godron’Del’Espece’tome2page52;
  Metzger’Syst。BeschreibungderKult。Kohlarten’1833s。6。)Icomparedtheflowersofalltheprincipalkinds;thoseoftheCouveTronchudaarewhiteandrathersmallerthanincommoncabbages;thoseofthePortsmouthbroccolihavenarrowersepals,andsmaller,lesselongatedpetals;andinnoothercabbagecouldanydifferencebedetected。Withrespecttotheseed—pods,inthepurpleKohlrabialone,dotheydiffer,beingalittlelongerandnarrowerthanusual。Imadeacollectionoftheseedsoftwenty—
  eightdifferentkinds,andmostofthemwereundistinguishable;whentherewasanydifferenceitwasexcessivelyslight;thus,theseedsofvariousbroccolisandcauliflowers,whenseeninmass,arealittleredder;thoseoftheearlygreenUlmsavoyarerathersmaller;andthoseoftheBredakailslightlylargerthanusual,butnotlargerthantheseedsofthewildcabbagefromthecoastofWales。Whatacontrastintheamountofdifferenceispresentedif,ontheonehand,wecomparetheleavesandstemsofthevariouskindsofcabbagewiththeirflowers,pods,andseeds,andontheotherhandthecorrespondingpartsinthevarietiesofmaizeandwheat!Theexplanationisobvious;theseedsalonearevaluedinourcereals,andtheirvariationshavebeenselected;whereastheseeds,seed—
  pods,andflowers,havebeenutterlyneglectedinthecabbage,whilstmanyusefulvariationsintheirleavesandstemshavebeennoticedandpreservedfromanextremelyremoteperiod,forcabbageswerecultivatedbytheoldCelts。(9/67。Regnier’Del’EconomiePubliquedesCeltes’1818page438。)
  Itwouldbeuselesstogiveaclassifieddescription(9/68。SeetheelderDeCandollein’Transact。ofHort。Soc。’volume5;andMetzger’Kohlarten’
  etc。)ofthenumerousraces,sub—races,andvarietiesofthecabbage;butitmaybementionedthatDr。Lindleyhaslatelyproposed(9/69。’Gardener’sChronicle’1859page992。)asystemfoundedonthestateofdevelopmentoftheterminalandlateralleaf—buds。Thus:
  I。Alltheleaf—budsactiveandopen,asinthewild—cabbage,kail,etc。
  II。Alltheleaf—budsactive,butformingheads,asinBrussel—sprouts,etc。
  III。Terminalleaf—budaloneactive,formingaheadasincommoncabbages,savoys,etc。
  IV。Terminalleaf—budaloneactive,andopen,withmostoftheflowersabortiveandsucculent,asinthecauliflowerandbroccoli。
  V。Alltheleaf—budsactiveandopen,withmostoftheflowersabortiveandsucculent,asinthesprouting—broccoli。Thislattervarietyisanewone,andbearsthesamerelationtocommonbroccoli,asBrussel—sproutsdotocommoncabbages;itsuddenlyappearedinabedofcommonbroccoli,andwasfoundfaithfullytotransmititsnewly—acquiredandremarkablecharacters。
  Theprincipalkindsofcabbageexistedatleastasearlyasthesixteenthcentury(9/70。Alph。DeCandolle’Geograph。Bot。’pages842and989。),sothatnumerousmodificationsofstructurehavebeeninheritedforalongperiod。Thisfactisthemoreremarkableasgreatcaremustbetakentopreventthecrossingofthedifferentkinds。Togiveproofofthis:I
  raised233seedlingsfromcabbagesofdifferentkinds,whichhadpurposelybeenplantedneareachother,andoftheseedlingsnolessthan155wereplainlydeterioratedandmongrelised;norweretheremaining78allperfectlytrue。Itmaybedoubtedwhethermanypermanentvarietieshavebeenformedbyintentionaloraccidentalcrosses;forsuchcrossedplantsarefoundtobeveryinconstant。Onekind,however,called"Cottager’sKail,"haslatelybeenproducedbycrossingcommonkailandBrussel—
  sprouts,recrossedwithpurplebroccoli(9/71。’Gardener’sChronicle’
  February1858page128。),andissaidtobetrue;butplantsraisedbymewerenotnearlysoconstantincharacterasanycommonkindofcabbage。
  Althoughmostofthekindskeeptrueifcarefullypreservedfromcrossing,yettheseed—bedsmustbeyearlyexamined,andafewseedlingsaregenerallyfoundfalse;buteveninthiscasetheforceofinheritanceisshown,for,asMetzgerhasremarked(9/72。’Kohlarten’s。22。)whenspeakingofBrussel—sprouts,thevariationsgenerallykeeptotheir"unterart,"ormainrace。Butinorderthatanykindmaybetrulypropagatedtheremustbenogreatchangeintheconditionsoflife;thuscabbageswillnotformheadsinhotcountries,andthesamethinghasbeenobservedwithanEnglishvarietygrownduringanextremelywarmanddampautumnnearParis。(9/73。Godron’Del’Espece’tome2page52;Metzger’Kohlarten’s。
  22。)Extremelypoorsoilalsoaffectsthecharactersofcertainvarieties。
  MostauthorsbelievethatalltheracesaredescendedfromthewildcabbagefoundonthewesternshoresofEurope;butAlph。DeCandolle(9/74。
  ’Geograph。Bot。’page840。)forciblyargues,onhistoricalandothergrounds,thatitismoreprobablethattwoorthreecloselyalliedforms,generallyrankedasdistinctspecies,stilllivingintheMediterraneanregion,aretheparents,nowallcommingledtogether,ofthevariouscultivatedkinds。Inthesamemanneraswehaveoftenseenwithdomesticatedanimals,thesupposedmultipleoriginofthecabbagethrowsnolightonthecharacteristicdifferencesbetweenthecultivatedforms。Ifourcabbagesarethedescendantsofthreeorfourdistinctspecies,everytraceofanysterilitywhichmayoriginallyhaveexistedbetweenthemisnowlost,fornoneofthevarietiescanbekeptdistinctwithoutscrupulouscaretopreventintercrossing。
  TheothercultivatedformsofthegenusBrassicaaredescended,accordingtotheviewadoptedbyGodronandMetzger(9/75。Godron’Del’Espece’tome2page54;Metzger’Kohlarten’s。10。),fromtwospecies,B。napusandrapa;butaccordingtootherbotanistsfromthreespecies;whilstothersagainstronglysuspectthatalltheseforms,bothwildandcultivated,oughttoberankedasasinglespecies。Brassicanapushasgivenrisetotwolargegroups,namely,Swedishturnips(believedtobeofhybridorigin)
  (9/76。’Gardener’sChronicleandAgricult。Gazette’1856page729。Seemoreespeciallyibid1868page275:thewriterassertsthatheplantedavarietyofcabbage(B。oleracea)closetoturnips(B。rapa)andraisedfromthecrossedseedlingstrueSwedishturnips。Theselatterplantsought,therefore,tobeclassedwithcabbagesorturnips,andnotunderB。napus。)
  andColzas,theseedsofwhichyieldoil。Brassicarapa(ofKoch)hasalsogivenrisetotworaces,namely,commonturnipsandtheoil—givingrape。
  Theevidenceisunusuallyclearthattheselatterplants,thoughsodifferentinexternalappearance,belongtothesamespecies;fortheturniphasbeenobservedbyKochandGodrontoloseitsthickrootsinuncultivatedsoil;andwhenrapeandturnipsaresowntogethertheycrosstosuchadegreethatscarcelyasingleplantcomestrue。(9/77。
  ’Gardener’sChronicleandAgricult。Gazette’1855page730。)Metzgerbycultureconvertedthebiennialorwinterrapeintotheannualorsummerrape,——varietieswhichhavebeenthoughtbysomeauthorstobespecificallydistinct。(9/78。Metzger,’Kohlarten’s。51。)
  Intheproductionoflarge,fleshy,turnip—likestems,wehaveacaseofanalogousvariationinthreeformswhicharegenerallyconsideredasdistinctspecies。Butscarcelyanymodificationseemssoeasilyacquiredasasucculentenlargementofthestemorroot——thatis,astoreofnutrimentlaidupfortheplant’sownfutureuse。Weseethisinourradishes,beet,andinthelessgenerallyknown"turnip—rooted"celery,andinthefinocchio,orItalianvarietyofthecommonfennel。Mr。Buckmanhaslatelyprovedbyhisinterestingexperimentsbowquicklytherootsofthewildparsnipcanbeenlarged,asVilmorinformerlyprovedinthecaseofthecarrot。(9/79。TheseexperimentsbyVilmorinhavebeenquotedbymanywriters。Aneminentbotanist,Prof。Decaisne,haslatelyexpresseddoubtsonthesubjectfromhisownnegativeresults,butthesecannotbevaluedequallywithpositiveresults。Ontheotherhand,M。Carrierehaslatelystated(’Gardener’sChronicle’1865page1154),thathetookseedfromawildcarrot,growingfarfromanycultivatedland,andeveninthefirstgenerationtherootsofhisseedlingsdifferedinbeingspindle—shaped,longer,softer,andlessfibrousthanthoseofthewildplant。Fromtheseseedlingsheraisedseveraldistinctvarieties。)
  Thislatterplant,initscultivatedstate,differsinscarcelyanycharacterfromthewildEnglishcarrot,exceptingeneralluxurianceandinthesizeandqualityofitsroots;buttenvarieties,differinginthecolour,shape,andqualityoftheroot,arecultivatedinEnglandandcometruebyseed。(9/80。Loudon’s’Encyclop。ofGardening’page835。)Hencewiththecarrot,asinsomanyothercases,forinstancewiththenumerousvarietiesandsub—varietiesoftheradish,thatpartoftheplantwhichisvaluedbyman,falselyappearsalonetohavevaried。Thetruthisthatvariationsinthispartalonehavebeenselected;andtheseedlingsinheritingatendencytovaryinthesameway,analogousmodificationshavebeenagainandagainselected,untilatlastagreatamountofchangehasbeeneffected。
  Withrespecttotheradish,M。Carriere,bysowingtheseedofthewildRaphanusraphanistruminrichsoil,andbycontinuedselectionduringseveralgenerations,raisedmanyvarieties,closelylikethecultivatedradish(R。sativus)intheirroots,aswellasthewonderfulChinesevariety,R。caudatus:(see’Journald’Agriculturepratique’tome11869
  page159;alsoaseparateessay’OriginedesPlantesDomestiques’1869。)