Indomitablepatience,thefinestpowersofdiscrimination,andsoundjudgmentmustbeexercisedduringmanyyears。Aclearlypredeterminedobjectmustbekeptsteadilyinview。Fewmenareendowedwithallthesequalities,especiallywiththatofdiscriminatingveryslightdifferences;judgmentcanbeacquiredonlybylongexperience;butifanyofthesequalitiesbewanting,thelabourofalifemaybethrownaway。Ihavebeenastonishedwhencelebratedbreeders,whoseskillandjudgmenthavebeenprovedbytheirsuccessatexhibitions,haveshownmetheiranimals,whichappearedallalike,andhaveassignedtheirreasonsformatchingthisandthatindividual。TheimportanceofthegreatprincipleofSelectionmainlyliesinthispowerofselectingscarcelyappreciabledifferences,whichneverthelessarefoundtobetransmissible,andwhichcanbeaccumulateduntiltheresultismademanifesttotheeyesofeverybeholder。
  Theprincipleofselectionmaybeconvenientlydividedintothreekinds。
  METHODICALSELECTIONisthatwhichguidesamanwhosystematicallyendeavourstomodifyabreedaccordingtosomepredeterminedstandard。UNCONSCIOUS
  SELECTIONisthatwhichfollowsfrommennaturallypreservingthemostvaluedanddestroyingthelessvaluedindividuals,withoutanythoughtofalteringthebreed;andundoubtedlythisprocessslowlyworksgreatchanges。
  Unconsciousselectiongraduatesintomethodical,andonlyextremecasescanbedistinctlyseparated;forhewhopreservesausefulorperfectanimalwillgenerallybreedfromitwiththehopeofgettingoffspringofthesamecharacter;butaslongashehasnotapredeterminedpurposetoimprovethebreed,hemaybesaidtobeselectingunconsciously。(20/1。Theterm"unconsciousselection"hasbeenobjectedtoasacontradiction;butseesomeexcellentobservationsonthisheadbyProf。Huxley(’Nat。Hist。Review’
  October1864page578)whoremarksthatwhenthewindheapsupsand—dunesitsiftsandUNCONSCIOUSLYSELECTSfromthegravelonthebeachgrainsofsandofequalsize。)Lastly,wehaveNATURALSELECTION,whichimpliesthattheindividualswhicharebestfittedforthecomplex,andinthecourseofageschangingconditionstowhichtheyareexposed,generallysurviveandprocreatetheirkind。Withdomesticproductions,naturalselectioncomestoacertainextentintoaction,independentlyof,andeveninoppositionto,thewillofman。
  METHODICALSELECTION。
  WhatmanhaseffectedwithinrecenttimesinEnglandbymethodicalselectionisclearlyshownbyourexhibitionsofimprovedquadrupedsandfancybirds。
  Withrespecttocattle,sheep,andpigs,weowetheirgreatimprovementtoalongseriesofwell—knownnames——Bakewell,Coiling,Ellman,Bates,JonasWebb,LordsLeicesterandWestern,FisherHobbs,andothers。Agriculturalwritersareunanimousonthepowerofselection:anynumberofstatementstothiseffectcouldbequoted;afewwillsuffice。Youatt,asagaciousandexperiencedobserver,writes(20/2。’OnSheep’1838page60。)theprincipleofselectionis"thatwhichenablestheagriculturist,notonlytomodifythecharacterofhisflock,buttochangeitaltogether。"AgreatbreederofShorthorns(20/3。Mr。J。WrightonShorthornCattlein’JournalofRoyalAgricult。Soc。’volume7pages208,209。)says,"IntheanatomyoftheshouldermodernbreedershavemadegreatimprovementontheKettonshorthornsbycorrectingthedefectintheknuckleorshoulder—joint,andbylayingthetopoftheshouldermoresnuglyinthecrop,andtherebyfillingupthehollowbehindit……Theeyehasitsfashionatdifferentperiods:atonetimetheeyehighandoutstandingfromthehead,andatanothertimethesleepyeyesunkintothehead;buttheseextremeshavemergedintothemediumofafull,clearandprominenteyewithaplacidlook。"
  Again,hearwhatanexcellentjudgeofpigs(20/4。H。D。Richardson’OnPigs’
  1847page44。)says:"Thelegsshouldbenolongerthanjusttopreventtheanimal’sbellyfromtrailingontheground。Thelegistheleastprofitableportionofthehog,andwethereforerequirenomoreofitthanisabsolutelynecessaryforthesupportoftherest。"Letanyonecomparethewild—boarwithanyimprovedbreed,andhewillseehoweffectuallythelegshavebeenshortened。
  Fewpersons,exceptbreeders,areawareofthesystematiccaretakeninselectinganimals,andofthenecessityofhavingaclearandalmostpropheticvisionintofuturity。LordSpencer’sskillandjudgmentwerewellknown;andhewrites(20/5。’JournalofRoyalAgricult。Soc。’volume1page24。),"Itisthereforeverydesirable,beforeanymancommencestobreedeithercattleorsheep,thatheshouldmakeuphismindtotheshapeandqualitieshewishestoobtain,andsteadilypursuethisobject。"LordSomerville,inspeakingofthemarvellousimprovementoftheNewLeicestersheep,effectedbyBakewellandhissuccessors,says,"Itwouldseemasiftheyhadfirstdrawnaperfectform,andthengivenitlife。"Youatt(20/6。’OnSheep’pages520,319。)urgesthenecessityofannuallydraftingeachflock,asmanyanimalswillcertainlydegenerate"fromthestandardofexcellencewhichthebreederhasestablishedinhisownmind。"Evenwithabirdofsuchlittleimportanceasthecanary,longago(1780—1790)ruleswereestablished,andastandardofperfectionwasfixedaccordingtowhichtheLondonfancierstriedtobreedtheseveralsub—
  varieties。(20/7。Loudon’s’Mag。ofNat。Hist。’volume81835page618。)A
  greatwinnerofprizesatthePigeon—shows(20/8。’AtreatiseontheArtofBreedingtheAlmondTumbler’1851page9。),indescribingtheshort—facedAlmondTumbler,says,"Therearemanyfirst—ratefancierswhoareparticularlypartialtowhatiscalledthegoldfinch—beak,whichisverybeautiful;otherssay,takeafull—sizeroundcherrythentakeabarleycorn,andjudiciouslyplacingandthrustingitintothecherry,formasitwereyourbeak;andthatisnotall,foritwillformagoodheadandbeak,provided,asIsaidbefore,itisjudiciouslydone;otherstakeanoat;butasIthinkthegoldfinch—beakthehandsomest,Iwouldadvisetheinexperiencedfanciertogettheheadofagoldfinch,andkeepitbyhimforhisobservation。"Wonderfullydifferentasarethebeaksoftherockpigeonandgoldfinch,theendhasundoubtedlybeennearlygained,asfarasexternalshapeandproportionsareconcerned。
  Notonlyshouldouranimalsbeexaminedwiththegreatestcarewhilstalive,but,asAndersonremarks(20/9。’RecreationsinAgriculture’volume2page409。)theircarcasesshouldbescrutinised,"soastobreedfromthedescendantsofsuchonlyas,inthelanguageofthebutcher,cutupwell。"The"grainofthemeat"incattle,anditsbeingwellmarbledwithfat(20/10。
  ’YouattonCattle’pages191,227。),andthegreaterorlessaccumulationoffatintheabdomenofoursheep,havebeenattendedtowithsuccess。Sowithpoultry,awriter(20/11。Ferguson’PrizePoultry’1854page208。),speakingofCochin—Chinafowls,whicharesaidtodiffermuchinthequalityoftheirflesh,says,"thebestmodeistopurchasetwoyoungbrother—cocks,kill,dress,andserveupone;ifhebeindifferent,similarlydisposeoftheother,andtryagain;if,however,hebefineandwell—flavoured,hisbrotherwillnotbeamissforbreedingpurposesforthetable。"
  Thegreatprincipleofthedivisionoflabourhasbeenbroughttobearonselection。Incertaindistricts(20/12。Wilsonin’Transact。HighlandAgricult。Soc。’quotedin’Gardener’sChronicle’1844page29。)"thebreedingofbullsisconfinedtoaverylimitednumberofpersons,whobydevotingtheirwholeattentiontothisdepartment,areablefromyeartoyeartofurnishaclassofbullswhicharesteadilyimprovingthegeneralbreedofthedistrict。"Therearingandlettingofchoiceramshaslongbeen,asiswellknown,achiefsourceofprofittoseveraleminentbreeders。InpartsofGermanythisprincipleiscarriedwithmerinosheeptoanextremepoint。
  (20/13。Simmondsquotedin’Gardener’sChronicle’1855page637。Andforthesecondquotationsee’YouattonSheep’page171。)So"importantistheproperselectionofbreedinganimalsconsidered,thatthebestflock—mastersdonottrusttotheirownjudgmentortothatoftheirshepherds,butemploypersonscalled’sheep—classifiers’whomakeittheirspecialbusinesstoattendtothispartofthemanagementofseveralflocks,andthustopreserve,orifpossibletoimprove,thebestqualitiesofbothparentsinthelambs。"InSaxony,"whenthelambsareweaned,eachinhisturnisplaceduponatablethathiswoolandformmaybeminutelyobserved。Thefinestareselectedforbreedingandreceiveafirstmark。Whentheyareoneyearold,andpriortoshearingthem,anothercloseexaminationofthosepreviouslymarkedtakesplace:thoseinwhichnodefectcanbefoundreceiveasecondmark,andtherestarecondemned。Afewmonthsafterwardsathirdandlastscrutinyismade;
  theprimeramsandewesreceiveathirdandfinalmark,buttheslightestblemishissufficienttocausetherejectionoftheanimal。"Thesesheeparebredandvaluedalmostexclusivelyforthefinenessoftheirwool;andtheresultcorrespondswiththelabourbestowedontheirselection。Instrumentshavebeeninventedtomeasureaccuratelythethicknessofthefibres;and"anAustrianfleecehasbeenproducedofwhichtwelvehairsequalledinthicknessonefromaLeicestersheep。"
  Throughouttheworld,whereversilkisproduced,thegreatestcareisbestowedonselectingthecocoonsfromwhichthemothsforbreedingaretobereared。A
  carefulcultivator(20/14。Robinet’VersaSoie’1848page271。)likewiseexaminesthemothsthemselves,anddestroysthosethatarenotperfect。ButwhatmoreimmediatelyconcernsusisthatcertainfamiliesinFrancedevotethemselvestoraisingeggsforsale。(20/15。Quatrefages’LesMaladiesduVeraSoie’1859page101。)InChina,nearShanghai,theinhabitantsoftwosmalldistrictshavetheprivilegeofraisingeggsforthewholesurroundingcountry,andthattheymaygiveuptheirwholetimetothisbusiness,theyareinterdictedbylawfromproducingsilk。(20/16。M。Simonin’Bull。delaSoc。
  d’Acclimat。’tome91862page221。)
  Thecarewhichsuccessfulbreederstakeinmatchingtheirbirdsissurprising。
  SirJohnSebright,whosefameisperpetuatedbythe"SebrightBantam,"usedtospend"twoandthreedaysinexamining,consulting,anddisputingwithafriendwhichwerethebestoffiveorsixbirds。"(20/17。’ThePoultryChronicle’volume11854page607。)Mr。Bult,whosepouter—pigeonswonsomanyprizes,andwereexportedtoNorthAmericaunderthechargeofamansentonpurpose,toldmethathealwaysdeliberatedforseveraldaysbeforehematchedeachpair。Hencewecanunderstandtheadviceofaneminentfancier,whowrites(20/18。J。M。Eaton’ATreatiseonFancyPigeons’1852page14and’A
  TreatiseontheAlmondTumbler’1851page11。)"Iwouldhereparticularlyguardyouagainsthavingtoogreatavarietyofpigeons,otherwiseyouwillknowalittleofall,butnothingaboutoneasitoughttobeknown。"
  Apparentlyittranscendsthepowerofthehumanintellecttobreedallkinds:
  "itispossiblethattheremaybeafewfanciersthathaveagoodgeneralknowledgeoffancypigeons;buttherearemanymorewholabourunderthedelusionofsupposingtheyknowwhattheydonot。"Theexcellenceofonesub—
  variety,theAlmondTumbler,liesintheplumage,carriage,head,beak,andeye;butitistoopresumptuousinthebeginnertotryforallthesepoints。
  Thegreatjudgeabovequotedsays,"Therearesomeyoungfancierswhoareover—covetous,whogoforalltheabovefivepropertiesatonce;theyhavetheirrewardbygettingnothing。"Wethusseethatbreedingevenfancypigeonsisnosimpleart:wemaysmileatthesolemnityoftheseprecepts,buthewholaughswillwinnoprizes。
  Whatmethodicalselectionhaseffectedforouranimalsissufficientlyproved,asalreadyremarked,byourExhibitions。Sogreatlywerethesheepbelongingtosomeoftheearlierbreeders,suchasBakewellandLordWestern,changed,thatmanypersonscouldnotbepersuadedthattheyhadnotbeencrossed。Ourpigs,asMr。Corringhamremarks(20/19。’JournalRoyalAgriculturalSoc。’
  volume6page22。)duringthelasttwentyyearshaveundergone,throughrigorousselectiontogetherwithcrossing,acompletemetamorphosis。ThefirstexhibitionforpoultrywasheldintheZoologicalGardensin1845;andtheimprovementeffectedsincethattimehasbeengreat。AsMr。Bailey,thegreatjudge,remarkedtome,itwasformerlyorderedthatthecomboftheSpanishcockshouldbeupright,andinfourorfiveyearsallgoodbirdshaduprightcombs;itwasorderedthatthePolishcockshouldhavenocomborwattles,andnowabirdthusfurnishedwouldbeatoncedisqualified;beardswereordered,andoutoffifty—sevenpenslately(1860)exhibitedattheCrystalPalace,allhadbeards。Soithasbeeninmanyothercases。Butinallcasesthejudgesorderonlywhatisoccasionallyproducedandwhatcanbeimprovedandrenderedconstantbyselection。Thesteadyincreaseinweightduringthelastfewyearsinourfowls,turkeys,ducks,andgeeseisnotorious;"six—poundducksarenowcommon,whereasfourpoundswasformerlytheaverage。"Asthetimerequiredtomakeachangehasnotoftenbeenrecorded,itmaybeworthmentioningthatittookMr。Wickingthirteenyearstoputacleanwhiteheadonanalmondtumbler’sbody,"atriumph,"saysanotherfancier,"ofwhichhemaybejustlyproud。"(20/20。’PoultryChronicle’volume21855page596。)
  Mr。Tollet,ofBetleyHall,selectedcows,andespeciallybulls,descendedfromgoodmilkers,forthesolepurposeofimprovinghiscattlefortheproductionofcheese;hesteadilytestedthemilkwiththelactometer,andineightyearsheincreased,asIwasinformedbyhim,theproductinproportionoffourtothree。Hereisacuriouscase(20/21。Isid。GeoffroySt。—Hilaire’Hist。Nat。Gen。’tome3page254。)ofsteadybutslowprogress,withtheendnotasyetfullyattained:in1784araceofsilkwormswasintroducedintoFrance,inwhichonehundredinthethousandfailedtoproducewhitecocoons;
  butnowaftercarefulselectionduringsixty—fivegenerations,theproportionofyellowcocoonshasbeenreducedtothirty—fiveinthethousand。
  Withplantsselectionhasbeenfollowedwiththesamegoodresultaswithanimals。Buttheprocessissimpler,forplantsinthegreatmajorityofcasesbearbothsexes。Nevertheless,withmostkindsitisnecessarytotakeasmuchcaretopreventcrossesaswithanimalsorunisexualplants;butwithsomeplants,suchaspeas,thiscareisnotnecessary。Withallimprovedplants,exceptingofcoursethosewhicharepropagatedbybuds,cuttings,etc。,itisalmostindispensabletoexaminetheseedlingsanddestroythosewhichdepartfromthepropertype。Thisiscalled"roguing,"andis,infact,aformofselection,liketherejectionofinferioranimals。Experiencedhorticulturistsandagriculturistsincessantlyurgeeveryonetopreservethefinestplantsfortheproductionofseed。
  Althoughplantsoftenpresentmuchmoreconspicuousvariationsthananimals,yettheclosestattentionisgenerallyrequisitetodetecteachslightandfavourablechange。Mr。Mastersrelates(20/22。’Gardener’sChronicle’1850
  page198。)how"manyapatienthourwasdevoted,"whilsthewasyoung,tothedetectionofdifferencesinpeasintendedforseed。Mr。Barnet(20/23。
  ’Transact。Hort。Soc。’volume6page152。)remarksthattheoldscarletAmericanstrawberrywascultivatedformorethanacenturywithoutproducingasinglevariety;andanotherwriterobserveshowsingularitwasthatwhengardenersfirstbegantoattendtothisfruititbegantovary;thetruthnodoubtbeingthatithadalwaysvaried,butthat,untilslightvariationswereselectedandpropagatedbyseed,noconspicuousresultwasobtained。Thefinestshadesofdifferenceinwheathavebeendiscriminatedandselectedwithalmostasmuchcareas,inthecaseofthehigheranimals,forinstancebyCol。LeCouteurandmoreespeciallybyMajorHallett。
  Itmaybeworthwhiletogiveafewexamplesofmethodicalselectionwithplants;butinfactthegreatimprovementofallourancientlycultivatedplantsmaybeattributedtoselectionlongcarriedon,inpartmethodically,andinpartunconsciously。Ihaveshowninaformerchapterhowtheweightofthegooseberryhasbeenincreasedbysystematicselectionandculture。TheflowersoftheHeartseasehavebeensimilarlyincreasedinsizeandregularityofoutline。WiththeCineraria,Mr。Glenny(20/24。’JournalofHorticulture’
  1862page369。)"wasboldenoughwhentheflowerswereraggedandstarryandilldefinedincolour,tofixastandardwhichwasthenconsideredoutrageouslyhighandimpossible,andwhich,evenifreached,itwassaid,weshouldbenogainersby,asitwouldspoilthebeautyoftheflowers。Hemaintainedthathewasright;andtheeventhasprovedittobeso。"Thedoublingofflowershasseveraltimesbeeneffectedbycarefulselection:theRev。W。Williamson(20/25’Transact。Hort。Soc。’volume4page381。),aftersowingduringseveralyearsseedofAnemonecoronaria,foundaplantwithoneadditionalpetal;hesowedtheseedofthis,andbyperseveranceinthesamecourseobtainedseveralvarietieswithsixorsevenrowsofpetals。ThesingleScotchrosewasdoubled,andyieldedeightgoodvarietiesinnineortenyears。(20/26。’Transact。Hort。Soc。’volume4page285。)TheCanterburybell(Campanulamedium)wasdoubledbycarefulselectioninfourgenerations。
  (20/27。Rev。W。Bromeheadin’Gardener’sChronicle’1857page550。)InfouryearsMr。Buckman(20/28。’Gardener’sChronicle’1862page721。),bycultureandcarefulselection,convertedparsnips,raisedfromwildseed,intoanewandgoodvariety。Byselectionduringalongcourseofyears,theearlymaturityofpeashasbeenhastenedbybetweentenandtwenty—onedays。(20/29。
  Dr。Andersonin’TheBee’volume6page96;Mr。Barnesin’Gardener’sChronicle’1844page476。)Amorecuriouscaseisofferedbythebeetplant,whichsinceitscultivationinFrance,hasalmostexactlydoubleditsyieldofsugar。Thishasbeeneffectedbythemostcarefulselection;thespecificgravityoftherootsbeingregularlytested,andthebestrootssavedfortheproductionofseed。(20/30。Godron’Del’Espece’1859tome2page69;
  ’Gardener’sChronicle’1854page258。)
  SELECTIONBYANCIENTANDSEMI—CIVILISEDPEOPLE。
  Inattributingsomuchimportancetotheselectionofanimalsandplants,itmaybeobjected,thatmethodicalselectionwouldnothavebeencarriedonduringancienttimes。Adistinguishednaturalistconsidersitasabsurdtosupposethatsemi—civilisedpeopleshouldhavepractisedselectionofanykind。Undoubtedlytheprinciplehasbeensystematicallyacknowledgedandfollowedtoafargreaterextentwithinthelasthundredyearsthanatanyformerperiod,andacorrespondingresulthasbeengained;butitwouldbeagreatererrortosuppose,asweshallimmediatelysee,thatitsimportancewasnotrecognisedandactedonduringthemostancienttimes,andbysemi—
  civilisedpeople。Ishouldpremisethatmanyfactsnowtobegivenonlyshowthatcarewastakeninbreeding;butwhenthisisthecase,selectionisalmostsuretobepractisedtoacertainextent。Weshallhereafterbeenabledbettertojudgehowfarselection,whenonlyoccasionallycarriedon,byafewoftheinhabitantsofacountry,willslowlyproduceagreateffect。
  Inawell—knownpassageinthethirtiethchapterofGenesis,rulesaregivenforinfluencing,aswasthenthoughtpossible,thecolourofsheep;andspeckledanddarkbreedsarespokenofasbeingkeptseparate。BythetimeofDavidthefleecewaslikenedtosnow。Youatt(20/31。’OnSheep’page18。),whohasdiscussedallthepassagesinrelationtobreedingintheOldTestament,concludesthatatthisearlyperiod"someofthebestprinciplesofbreedingmusthavebeensteadilyandlongpursued。"Itwasordered,accordingtoMoses,that"Thoushaltnotletthycattlegenderwithadiversekind;"butmuleswerepurchased(20/32。Volz’BeitragezurKulturgeschichte’1852s。47。)sothatatthisearlyperiodothernationsmusthavecrossedthehorseandass。
  Itissaid(20/33。Mitford’HistoryofGreece’volume1page73。)thatErichthonius,somegenerationsbeforetheTrojanwar,hadmanybrood—mares,"whichbyhiscareandjudgmentinthechoiceofstallionsproducedabreedofhorsessuperiortoanyinthesurroundingcountries。"Homer(Book5)speaksofAeneas’horsesasbredfrommareswhichwereputtothesteedsofLaomedon。
  Plato,inhis’Republic’saystoGlaucus,"Iseethatyouraiseatyourhouseagreatmanydogsforthechase。Doyoutakecareaboutbreedingandpairingthem?Amonganimalsofgoodblood,aretherenotalwayssomewhicharesuperiortotherest?"TowhichGlaucusanswersintheaffirmative。(20/34。
  Dr。Dallytranslatedin’AnthropologicalReview’May1864page101。)AlexandertheGreatselectedthefinestIndiancattletosendtoMacedoniatoimprovethebreed。(20/35。Volz’Beitrage’etc。1852s。80。)AccordingtoPliny(20/36
  ’HistoryoftheWorld’chapter45。),KingPyrrhushadanespeciallyvaluablebreedofoxen:andhedidnotsufferthebullsandcowstocometogethertillfouryearsold,thatthebreedmightnotdegenerate。Virgil,inhisGeorgics(lib。3),givesasstrongadviceasanymodernagriculturistcoulddo,carefullytoselectthebreedingstock;"tonotethetribe,thelineage,andthesire;whomtoreserveforhusbandoftheherd;"——tobrandtheprogeny;——toselectsheepofthepurestwhite,andtoexamineiftheirtonguesareswarthy。
  WehaveseenthattheRomanskeptpedigreesoftheirpigeons,andthiswouldhavebeenasenselessproceedinghadnotgreatcarebeentakeninbreedingthem。Columellagivesdetailedinstructionsaboutbreedingfowls:"Letthebreedinghensthereforebeofachoicecolour,arobustbody,square—built,full—breasted,withlargeheads,withuprightandbright—redcombs。Thosearebelievedtobethebestbredwhichhavefivetoes。"(20/37。’Gardener’sChronicle’1848page323。)AccordingtoTacitus,theCeltsattendedtotheracesoftheirdomesticanimals;andCaesarstatesthattheypaidhighpricestomerchantsforfineimportedhorses。(20/38。Reynier’Del’EconomiedesCeltes’1818pages487,503。)Inregardtoplants,Virgilspeaksofyearlycullingthelargestseeds;andCelsussays,"wherethecornandcropisbutsmall,wemustpickoutthebestearsofcorn,andofthemlayupourseedseparatelybyitself。"(20/39。LeCouteuron’Wheat’page15。)
  Comingdownthestreamoftime,wemaybebrief。AtaboutthebeginningoftheninthcenturyCharlemagneexpresslyorderedhisofficerstotakegreatcareofhisstallions;andifanyprovedbadorold,toforewarnhimingoodtimebeforetheywereputtothemares。(20/40。Michel’DesHaras’1861page84。)
  EveninacountrysolittlecivilisedasIrelandduringtheninthcentury,itwouldappearfromsomeancientverses(20/41。SirW。Wildean’EssayonUnmanufacturedAnimalRemains’etc。1860page11。),describingaransomdemandedbyCormac,thatanimalsfromparticularplaces,orhavingaparticularcharacter,werevalued。Thusitissaid,——
  TwopigsofthepigsofMacLir,Aramandewebothroundandred,IbroughtwithmefromAengus。
  IbroughtwithmeastallionandamareFromthebeautifulstudofManannan,AbullandawhitecowfromDruimCain。
  Athelstan,in930,receivedrunning—horsesasapresentfromGermany;andheprohibitedtheexportationofEnglishhorses。KingJohnimported"onehundredchosenstallionsfromFlanders。"(20/42。Col。HamiltonSmith’Nat。Library’
  volume12Horses,pages135,140。)OnJune16th,1305,thePrinceofWaleswrotetotheArchbishopofCanterbury,beggingfortheloanofanychoicestallion,andpromisingitsreturnattheendoftheseason。(20/43。Michel’DesHaras’page90。)TherearenumerousrecordsatancientperiodsinEnglishhistoryoftheimportationofchoiceanimalsofvariouskinds,andoffoolishlawsagainsttheirexportation。InthereignsofHenryVII。andVIII。itwasorderedthatthemagistrates,atMichaelmas,shouldscourtheheathsandcommons,anddestroyallmaresbeneathacertainsize。(20/44。Mr。Baker’HistoryoftheHorse’’Veterinary’volume13page423。)Someofourearlierkingspassedlawsagainsttheslaughteringramsofanygoodbreedbeforetheyweresevenyearsold,sothattheymighthavetimetobreed。InSpainCardinalXimenesissued,in1509,regulationsontheSELECTIONofgoodramsforbreeding。(20/45。M。l’AbbeCarlierin’JournaldePhysique’volume241784
  page181;thismemoircontainsmuchinformationontheancientselectionofsheep;andismyauthorityforramsnotbeingkilledyounginEngland。)
  TheEmperorAkbarKhanbeforetheyearl600issaidtohave"wonderfullyimproved"hispigeonsbycrossingthebreeds;andthisnecessarilyimpliescarefulselection。AboutthesameperiodtheDutchattendedwiththegreatestcaretothebreedingofthesebirds。Belonin1555saysthatgoodmanagersinFranceexaminedthecolouroftheirgoslingsinordertogetgeeseofawhitecolourandbetterkinds。Markhamin1631tellsthebreeder"toelectthelargestandgoodliestconies,"andentersintominutedetails。Evenwithrespecttoseedsofplantsfortheflower—garden,SirJ。Hanmerwritingabouttheyear1660(20/46。’Gardener’sChronicle’1843page389。)says,in"choosingseed,thebestseedisthemostweighty,andishadfromthelustiestandmostvigorousstems;"andhethengivesrulesaboutleavingonlyafewflowersonplantsforseed;sothatevensuchdetailswereattendedtoinourflower—gardenstwohundredyearsago。Inordertoshowthatselectionhasbeensilentlycarriedoninplaceswhereitwouldnothavebeenexpected,Imayaddthatinthemiddleofthelastcentury,inaremotepartofNorthAmerica,Mr。Cooperimprovedbycarefulselectionallhisvegetables,"sothattheyweregreatlysuperiortothoseofanyotherperson。Whenhisradishes,forinstance,arefitforuse,hetakestenortwelvethathemostapproves,andplantsthematleast100yardsfromothersthatblossomatthesametime。
  Inthesamemannerhetreatsallhisotherplants,varyingthecircumstancesaccordingtotheirnature。"(20/47。’CommunicationstoBoardofAgriculture’
  quotedinDr。Darwin’Phytologia’1800page451。)
  InthegreatworkonChinapublishedinthelastcenturybytheJesuits,andwhichischieflycompiledfromancientChineseencyclopaedias,itissaidthatwithsheep"improvingthebreedconsistsinchoosingwithparticularcarethelambswhicharedestinedforpropagation,innourishingthemwell,andinkeepingtheflocksseparate。"ThesameprincipleswereappliedbytheChinesetovariousplantsandfruit—trees。(20/48。’MemoiresurlesChinois’1786tome11page55;tome5page507。)Animperialedictrecommendsthechoiceofseedofremarkablesize;andselectionwaspractisedevenbyimperialhands,foritissaidthattheYa—mi,orimperialrice,wasnoticedatanancientperiodinafieldbytheEmperorKhang—hi,wassavedandcultivatedinhisgarden,andhassincebecomevaluablefrombeingtheonlykindwhichwillgrownorthoftheGreatWall。(20/49。’Recherchessurl’AgriculturedesChinois’parL。
  D’HerveySaint—Denys1850page229。WithrespecttoKhang—hiseeHuc’s’ChineseEmpire’page311。)Evenwithflowers,thetreepaeony(P。moutan)hasbeencultivated,accordingtoChinesetraditions,for1400years;between200
  and300varietieshavebeenraised,whicharecherishedliketulipsformerlywerebytheDutch。(20/50。Andersonin’Linn。Transact。’volume12page253。)
  Turningnowtosemi—civilisedpeopleandtosavages:itoccurredtome,fromwhatIhadseenofseveralpartsofSouthAmerica,wherefencesdonotexist,andwheretheanimalsareoflittlevalue,thattherewouldbeabsolutelynocareinbreedingorselectingthem;andthistoalargeextentistrue。Roulin(20/51。’Mem。del’Acad。’(diverssavants),tome61835page333。),however,describesinColumbiaanakedraceofcattle,whicharenotallowedtoincrease,onaccountoftheirdelicateconstitution。AccordingtoAzara(20/52。’DesQuadrupedesduParaguay’1801tome2pages333,371。)horsesareoftenborninParaguaywithcurlyhair;but,asthenativesdonotlikethem,theyaredestroyed。Ontheotherhand,Azarastatesthatahornlessbull,bornin1770,waspreservedandpropagateditsrace。IwasinformedoftheexistenceinBandaOrientalofabreedwithreversedhair;andtheextraordinaryniatacattlefirstappearedandhavesincebeenkeptdistinctinLaPlata。Hencecertainconspicuousvariationshavebeenpreserved,andothershavebeenhabituallydestroyed,inthesecountries,whicharesolittlefavourableforcarefulselection。Wehavealsoseenthattheinhabitantssometimesintroducefreshcattleontheirestatestopreventtheevileffectsofcloseinterbreeding。Ontheotherhand,IhaveheardonreliableauthoritythattheGauchosofthePampasnevertakeanypainsinselectingthebestbullsorstallionsforbreeding;andthisprobablyaccountsforthecattleandhorsesbeingremarkablyuniformincharacterthroughouttheimmenserangeoftheArgentinerepublic。
  LookingtotheOldWorld,intheSaharaDesert"TheTouaregisascarefulintheselectionofhisbreedingMahari(afineraceofthedromedary)astheArabisinthatofhishorse。Thepedigreesarehandeddown,andmanyadromedarycanboastagenealogyfarlongerthanthedescendantsoftheDarleyArabian。"(20/53。’TheGreatSahara’bytheRev。H。B。Tristram1860page238。)
  AccordingtoPallastheMongoliansendeavourtobreedtheYaksorhorse—tailedbuffaloeswithwhitetails,forthesearesoldtotheChinesemandarinsasfly—flappers;andMoorcroft,aboutseventyyearsafterPallas,foundthatwhite—tailedanimalswerestillselectedforbreeding。(20/54。Pallas’Act。
  Acad。St。Petersburg’1777page249。MoorcroftandTrebeck’TravelsintheHimalayanProvinces’1841。)
  WehaveseeninthechapterontheDogthatsavagesindifferentpartsofNorthAmericaandinGuianacrosstheirdogswithwildCanidae,asdidtheancientGauls,accordingtoPliny。Thiswasdonetogivetheirdogsstrengthandvigour,inthesamewayasthekeepersinlargewarrensnowsometimescrosstheirferrets(asIhavebeeninformedbyMr。Yarrell)withthewildpolecat,"togivethemmoredevil。"AccordingtoVarro,thewildasswasformerlycaughtandcrossedwiththetameanimaltoimprovethebreed,inthesamemannerasatthepresentdaythenativesofJavasometimesdrivetheircattleintotheforeststocrosswiththewildBanteng(Bossondaicus)。
  (20/55。QuotedfromRafflesinthe’IndianField’1859page196:forVarroseePallasutsupra。)InNorthernSiberia,amongtheOstyaks,thedogsvaryinmarkingsindifferentdistricts,butineachplacetheyarespottedblackandwhiteinaremarkablyuniformmanner(20/56。Erman’TravelsinSiberia’
  Englishtranslationvolume1page453。);andfromthisfactalonewemayinfercarefulbreeding,moreespeciallyasthedogsofonelocalityarefamedthroughoutthecountryfortheirsuperiority。IhaveheardofcertaintribesofEsquimauxwhotakeprideintheirteamsofdogsbeinguniformlycoloured。
  InGuiana,asSirH。Schomburgkinformsme(20/57。Seealso’JournalofR。
  Geograph。Soc。’volume13part1page65。),thedogsoftheTurumaIndiansarehighlyvaluedandextensivelybartered:thepriceofagoodoneisthesameasthatgivenforawife:theyarekeptinasortofcage,andtheIndians"takegreatcarewhenthefemaleisinseasontopreventherunitingwithadogofaninferiordescription。"TheIndianstoldSirRobertthat,ifadogprovedbadoruseless,hewasnotkilled,butwaslefttodiefromsheerneglect。
  HardlyanynationismorebarbarousthantheFuegians,butIhearfromMr。
  Bridges,theCatechisttotheMission,that,"whenthesesavageshavealarge,strong,andactivebitch,theytakecaretoputhertoafinedog,andeventakecaretofeedherwell,thatheryoungmaybestrongandwellfavoured。"
  IntheinteriorofAfrica,negroes,whohavenotassociatedwithwhitemen,showgreatanxietytoimprovetheiranimals;they"alwayschoosethelargerandstrongermalesforstock;"theMalakoloweremuchpleasedatLivingstone’spromisetosendthemabull,andsomeBakalolocarriedalivecockallthewayfromLoandaintotheinterior。(20/58。Livingstone’FirstTravels’pages191,439,565;seealso’ExpeditiontotheZambesi’1865page495,forananalogouscaserespectingagoodbreedofgoats。)AtFalabaMr。WinwoodReadenoticedanunusuallyfinehorse,andthenegroKinginformedhimthat"theownerwasnotedforhisskillinbreedinghorses。"Furthersouthonthesamecontinent,AnderssonstatesthathehasknownaDamaragivetwofineoxenforadogwhichstruckhisfancy。TheDamarastakegreatdelightinhavingwholedrovesofcattleofthesamecolour,andtheyprizetheiroxeninproportiontothesizeoftheirhorns。"TheNamaquashaveaperfectmaniaforauniformteam;andalmostallthepeopleofSouthernAfricavaluetheircattlenexttotheirwomen,andtakeaprideinpossessinganimalsthatlookhigh—bred。Theyrarelyornevermakeuseofahandsomeanimalasabeastofburden。"(20/59。
  Andersson’TravelsinSouthAfrica’pages232,318,319。)Thepowerofdiscriminationwhichthesesavagespossessiswonderful,andtheycanrecognisetowhichtribeanycattlebelong。Mr。Anderssonfurtherinformsmethatthenativesfrequentlymatchaparticularbullwithaparticularcow。
  Themostcuriouscaseofselectionbysemi—civilisedpeople,orindeedbyanypeople,whichIhavefoundrecorded,isthatgivenbyGarcilazodelaVega,adescendantoftheIncas,ashavingbeenpractisedinPerubeforethecountrywassubjugatedbytheSpaniards。(20/60。Dr。Vavasseurin’Bull。deLaSoc。
  d’Acclimat。’tome81861page136。)TheIncasannuallyheldgreathunts,whenallthewildanimalsweredrivenfromanimmensecircuittoacentralpoint。
  Thebeastsofpreywerefirstdestroyedasinjurious。ThewildGuanacosandVicunasweresheared;theoldmalesandfemaleskilled,andtheotherssetatliberty。Thevariouskindsofdeerwereexamined;theoldmalesandfemaleswerelikewisekilled,"buttheyoungfemales,withacertainnumberofmales,selectedfromthemostbeautifulandstrong,"weregiventheirfreedom。Here,then,wehaveselectionbymanaidingnaturalselection。SothattheIncasfollowedexactlythereversesystemofthatwhichourScottishsportsmanareaccusedoffollowing,namely,ofsteadilykillingthefineststags,thuscausingthewholeracetodegenerate。(20/61。’TheNaturalHistoryofDeeSide’1855page476。)Inregardtothedomesticatedllamasandalpacas,theywereseparatedinthetimeoftheIncasaccordingtocolour:andifbychanceoneinaflockwasbornofthewrongcolour,itwaseventuallyputintoanotherflock。
  InthegenusAucheniatherearefourforms,——theGuanacoandVicuna,foundwildandundoubtedlydistinctspecies;theLlamaandAlpaca,knownonlyinadomesticatedcondition。Thesefouranimalsappearsodifferent,thatmostnaturalists,especiallythosewhohavestudiedtheseanimalsintheirnativecountry,maintainthattheyarespecificallydistinct,notwithstandingthatnoonepretendstohaveseenawildllamaoralpaca。Mr。Ledger,however,whohascloselystudiedtheseanimalsbothinPeruandduringtheirexportationtoAustralia,andwhohasmademanyexperimentsontheirpropagation,adducesarguments(20/62。’Bull。delaSoc。d’Acclimat。’tome71860page457。)whichseemtomeconclusive,thatthellamaisthedomesticateddescendantoftheguanaco,andthealpacaofthevicuna。Andnowthatweknowthattheseanimalsweresystematicallybredandselectedmanycenturiesago,thereisnothingsurprisinginthegreatamountofchangewhichtheyhaveundergone。
  Itappearedtomeatonetimeprobablethat,thoughancientandsemi—civilisedpeoplemighthaveattendedtotheimprovementoftheirmoreusefulanimalsinessentialpoints,yetthattheywouldhavedisregardedunimportantcharacters。
  Buthumannatureisthesamethroughouttheworld:fashioneverywherereignssupreme,andmanisapttovaluewhateverhemaychancetopossess。WehaveseenthatinSouthAmericatheniatacattle,whichcertainlyarenotmadeusefulbytheirshortenedfacesandupturnednostrils,havebeenpreserved。
  TheDamarasofSouthAfricavaluetheircattleforuniformityofcolourandenormouslylonghorns。AndIwillnowshowthatthereishardlyanypeculiarityinourmostusefulanimalswhich,fromfashion,superstition,orsomeothermotive,hasnotbeenvalued,andconsequentlypreserved。Withrespecttocattle,"anearlyrecord,"accordingtoYouatt(20/63。’Cattle’
  page48。)"speaksofahundredwhitecowswithredearsbeingdemandedasacompensationbytheprincesofNorthandSouthWales。Ifthecattlewereofadarkorblackcolour,150weretobepresented。"SothatcolourwasattendedtoinWalesbeforeitssubjugationbyEngland。InCentralAfrica,anoxthatbeatsthegroundwithitstailiskilled;andinSouthAfricasomeoftheDamaraswillnoteatthefleshofaspottedox。TheKaffirsvalueananimalwithamusicalvoice;and"atasaleinBritishKaffrariathelow"ofaheiferexcitedsomuchadmirationthatasharpcompetitionsprungupforherpossession,andsherealisedaconsiderableprice。"(20/64。Livingstone’Travels’page576;Andersson’LakeNgami’1856page222。WithrespecttothesaleinKaffrariasee’QuarterlyReview’1860page139。)Withrespecttosheep,theChinesepreferramswithouthorns;theTartarspreferthemwithspirallywoundhorns,becausethehornlessarethoughttolosecourage。
  (20/65。’MemoiresurlesChinois’bytheJesuits1786tome11page57。)SomeoftheDamaraswillnoteatthefleshofhornlesssheep。Inregardtohorses,attheendofthefifteenthcenturyanimalsofthecolourdescribedasliartpommeweremostvaluedinFrance。TheArabshaveaproverb,"Neverbuyahorsewithfourwhitefeet,forhecarrieshisshroudwithhim"(20/66。F。Michel’DesHaras’pages47,50。);theArabsalso,aswehaveseen,despisedun—
  colouredhorses。Sowithdogs,Xenophonandothersatanancientperiodwereprejudicedinfavourofcertaincolours;and"whiteorslate—colouredhuntingdogswerenotesteemed。"(20/67。Col。HamiltonSmith’Dogs’in’Nat。Lib。’
  volume10page103。)
  Turningtopoultry,theoldRomangourmandsthoughtthattheliverofawhitegoosewasthemostsavoury。InParaguayblack—skinnedfowlsarekeptbecausetheyarethoughttobemoreproductive,andtheirfleshthemostproperforinvalids。(20/68。Azara’QuadrupedesduParaguay’tome2page324。)InGuiana,asIaminformedbySirR。Schomburgk,theaborigineswillnoteatthefleshoreggsofthefowl,buttworacesarekeptdistinctmerelyforornament。InthePhilippines,nolessthanninesub—varietiesofthegame—cockarekeptandnamed,sothattheymustbeseparatelybred。
  AtthepresenttimeinEurope,thesmallestpeculiaritiesarecarefullyattendedtoinourmostusefulanimals,eitherfromfashion,orasamarkofpurityofblood。Manyexamplescouldbegiven;twowillsuffice。"IntheWesterncountiesofEnglandtheprejudiceagainstawhitepigisnearlyasstrongasagainstablackoneinYorkshire。"InoneoftheBerkshiresub—
  breeds,itissaid,"thewhiteshouldbeconfinedtofourwhitefeet,awhitespotbetweentheeyes,andafewwhitehairsbehindeachshoulder。"Mr。
  Saddlerpossessedthreehundredpigs,everyoneofwhichwasmarkedinthismanner。"(20/69。Sidney’seditionofYouatt1860pages24,25。)Marshall,towardsthecloseofthelastcentury,inspeakingofachangeinoneoftheYorkshirebreedsofcattle,saysthehornshavebeenconsiderablymodified,as"aclean,small,sharphornhasbeenFASHIONABLEforthelasttwentyyears。"
  (20/70。’RuralEconomyofYorkshire’volume2page182。)InapartofGermanythecattleoftheRacedeGfoehlarevaluedformanygoodqualities,buttheymusthavehornsofaparticularcurvatureandtint,somuchsothatmechanicalmeansareappliediftheytakeawrongdirection;buttheinhabitants"consideritofthehighestimportancethatthenostrilsofthebullshouldbeflesh—coloured,andtheeyelasheslight;thisisanindispensablecondition。A
  calfwithbluenostrilswouldnotbepurchased,orpurchasedataverylowprice。"(20/71。MolletGayot’DuBoeuf’1860page547。)Thereforeletnomansaythatanypointorcharacteristootriflingtobemethodicallyattendedtoandselectedbybreeders。
  UNCONSCIOUSSELECTION。
  BythistermImean,asalreadymorethanonceexplained,thepreservationbymanofthemostvalued,andthedestructionoftheleastvaluedindividuals,withoutanyconsciousintentiononhispartofalteringthebreed。Itisdifficulttoofferdirectproofsoftheresultswhichfollowfromthiskindofselection;buttheindirectevidenceisabundant。Infact,exceptthatintheonecasemanactsintentionally,andintheotherunintentionally,thereislittledifferencebetweenmethodicalandunconsciousselection。Inbothcasesmanpreservestheanimalswhicharemostusefulorpleasingtohim,anddestroysorneglectstheothers。Butnodoubtafarmorerapidresultfollowsfrommethodicalthanfromunconsciousselection。The"roguing"ofplantsbygardeners,andthedestructionbylawinHenryVIII。’sreignofallunder—
  sizedmares,areinstancesofaprocessthereverseofselectionintheordinarysenseoftheword,butleadingtothesamegeneralresult。Theinfluenceofthedestructionofindividualshavingaparticularcharacteriswellshownbythenecessityofkillingeverylambwithatraceofblackaboutit,inordertokeeptheflockwhite;oragain,bytheeffectsontheaverageheightofthemenofFranceofthedestructivewarsofNapoleon,bywhichmanytallmenwerekilled,theshortonesbeinglefttobethefathersoffamilies。
  Thisatleastistheconclusionofsomeofthosewhohavecloselystudiedtheeffectsoftheconscription;anditiscertainthatsinceNapoleon’stimethestandardforthearmyhasbeenloweredtwoorthreetimes。
  Unconsciousselectionblendswithmethodical,sothatitisscarcelypossibletoseparatethem。Whenafancierlongagofirsthappenedtonoticeapigeonwithanunusuallyshortbeak,oronewiththetail—feathersunusuallydeveloped,althoughhebredfromthesebirdswiththedistinctintentionofpropagatingthevariety,yethecouldnothaveintendedtomakeashort—facedtumblerorafantail,andwasfarfromknowingthathehadmadethefirststeptowardsthisend。Ifhecouldhaveseenthefinalresult,hewouldhavebeenstruckwithastonishment,but,fromwhatweknowofthehabitsoffanciers,probablynotwithadmiration。OurEnglishcarriers,barbs,andshort—facedtumblershavebeengreatlymodifiedinthesamemanner,aswemayinferbothfromthehistoricalevidencegiveninthechaptersonthePigeon,andfromthecomparisonofbirdsbroughtfromdistantcountries。
  Soithasbeenwithdogs;ourpresentfox—houndsdifferfromtheoldEnglishhound;ourgreyhoundshavebecomelighter:theScotchdeer—houndhasbeenmodified,andisnowrare。Ourbulldogsdifferfromthosewhichwereformerlyusedforbaitingbulls。OurpointersandNewfoundlandsdonotcloselyresembleanynativedognowfoundinthecountrieswhencetheywerebrought。Thesechangeshavebeeneffectedpartlybycrosses;butineverycasetheresulthasbeengovernedbythestrictestselection。Nevertheless,thereisnoreasontosupposethatmanintentionallyandmethodicallymadethebreedsexactlywhattheynoware。Asourhorsesbecamefleeter,andthecountrymorecultivatedandsmoother,fleeterfox—houndsweredesiredandproduced,butprobablywithoutanyonedistinctlyforeseeingwhattheywouldbecome。Ourpointersandsetters,thelatteralmostcertainlydescendedfromlargespaniels,havebeengreatlymodifiedinaccordancewithfashionandthedesireforincreasedspeed。Wolveshavebecomeextinct,andsohasthewolf—dog;deerhavebecomerarer,bullsarenolongerbaited,andthecorrespondingbreedsofthedoghaveansweredtothechange。Butwemayfeelalmostsurethatwhen,forinstance,bullswerenolongerbaited,nomansaidtohimself,Iwillnowbreedmydogsofsmallersize,andthuscreatethepresentrace。Ascircumstanceschanged,menunconsciouslyandslowlymodifiedtheircourseofselection。
  Withracehorsesselectionforswiftnesshasbeenfollowedmethodically,andourhorsesnoweasilysurpasstheirprogenitors。TheincreasedsizeanddifferentappearanceoftheEnglishracehorseledagoodobserverinIndiatoask,"Couldanyoneinthisyearof1856,lookingatourracehorses,conceivethattheyweretheresultoftheunionoftheArabhorseandtheAfricanmare?"(20/72。’TheIndiaSportingReview’volume2page181;’TheStudFarm’
  byCecilpage58。)Thischangehas,itisprobable,beenlargelyeffectedthroughunconsciousselection,thatis,bythegeneralwishtobreedasfinehorsesaspossibleineachgeneration,combinedwithtrainingandhighfeeding,butwithoutanyintentiontogivetothemtheirpresentappearance。
  AccordingtoYouatt(20/73。’TheHorse’page22。),theintroductioninOliverCromwell’stimeofthreecelebratedEasternstallionsspeedilyaffectedtheEnglishbreed;"sothatLordHarleigh,oneoftheoldschool,complainedthatthegreathorsewasfastdisappearing。"Thisisanexcellentproofhowcarefullyselectionmusthavebeenattendedto;forwithoutsuchcare,alltracesofsosmallaninfusionofEasternbloodwouldsoonhavebeenabsorbedandlost。NotwithstandingthattheclimateofEnglandhasneverbeenesteemedparticularlyfavourabletothehorse,yetlong—continuedselection,bothmethodicalandunconscious,togetherwiththatpractisedbytheArabsduringastilllongerandearlierperiod,hasendedingivingusthebestbreedofhorsesintheworld。Macaulay(20/74。’HistoryofEngland’volume1page316。)
  remarks,"Twomenwhoseauthorityonsuchsubjectswasheldingreatesteem,theDukeofNewcastleandSirJohnFenwick,pronouncedthatthemeanesthackeverimportedfromTangierwouldproduceafinerprogenythancouldbeexpectedfromthebestsireofournativebreed。TheywouldnotreadilyhavebelievedthatatimewouldcomewhentheprincesandnoblesofneighbouringlandswouldbeaseagertoobtainhorsesfromEnglandasevertheEnglishhadbeentoobtainhorsesfromBarbary。"
  TheLondondray—horse,whichdifferssomuchinappearancefromanynaturalspecies,andwhichfromitssizehassoastonishedmanyEasternprinces,wasprobablyformedbytheheaviestandmostpowerfulanimalshavingbeenselectedduringmanygenerationsinFlandersandEngland,butwithouttheleastintentionorexpectationofcreatingahorsesuchaswenowsee。Ifwegobacktoanearlyperiodofhistory,webeholdintheantiqueGreekstatues,asSchaaffhausenhasremarked(20/75。’UeberBestandigkeitderArten。’),ahorseequallyunlikearaceordrayhorse,anddifferingfromanyexistingbreed。
  Theresultsofunconsciousselection,inanearlystage,arewellshowninthedifferencebetweentheflocksdescendedfromthesamestock,butseparatelyrearedbycarefulbreeders。YouattgivesanexcellentinstanceofthisfactinthesheepbelongingtoMessrs。BuckleyandBurgess,which"havebeenpurelybredfromtheoriginalstockofMr。Bakewellforupwardsoffiftyyears。ThereisnotasuspicionexistinginthemindofanyoneatallacquaintedwiththesubjectthattheownerofeitherflockhasdeviatedinanyoneinstancefromthepurebloodofMr。Bakewell’sflock;yetthedifferencebetweenthesheeppossessedbythesetwogentlemenissogreat,thattheyhavetheappearanceofbeingquitedifferentvarieties。"(20/76。’YouattonSheep’page315。)Ihaveseenseveralanalogousandwellmarkedcaseswithpigeons:forinstance,IhadafamilyofbarbsdescendedfromthoselongbredbySirJ。Sebright,andanotherfamilylongbredbyanotherfancier,andthetwofamiliesplainlydifferedfromeachother。Nathusius——andamorecompetentwitnesscouldnotbecited——observesthat,thoughtheShorthornsareremarkablyuniforminappearance(exceptincolour),yettheindividualcharacterandwishesofeachbreederbecomeimpressedonhiscattle,sothatdifferentherdsdifferslightlyfromoneanother。(20/77。’UeberShorthornRindvieh’1857s。51。)TheHerefordcattleassumedtheirpresentwell—markedcharactersoonaftertheyear1769,throughcarefulselectionbyMr。Tomkins(20/78。Low’DomesticatedAnimals’1845page363。)andthebreedhaslatelysplitintotwostrains——onestrainhavingawhiteface,anddifferingslightly,itissaid(20/79。