Ofterrestrialmammals,thereisonlyonewhichmustbeconsideredasindigenous,namely,amouseMusGalapagoensis,andthisisconfined,asfarasIcouldascertain,toChathamIsland,themosteasterlyislandofthegroup。Itbelongs,asIaminformedbyMr。Waterhouse,toadivisionofthefamilyofmicecharacteristicofAmerica。AtJamesIsland,thereisaratsufficientlydistinctfromthecommonkindtohavebeennamedanddescribedbyMr。Waterhouse;butasitbelongstotheold-worlddivisionofthefamily,andasthisislandhasbeenfrequentedbyshipsforthelasthundredandfiftyyears,Icanhardlydoubtthatthisratismerelyavarietyproducedbythenewandpeculiarclimate,food,andsoil,towhichithasbeensubjected。Althoughnoonehasarighttospeculatewithoutdistinctfacts,yetevenwithrespecttotheChathamIslandmouse,itshouldbeborneinmind,thatitmaypossiblybeanAmericanspeciesimportedhere;forIhaveseen,inamostunfrequentedpartofthePampas,anativemouselivingintheroofofanewlybuilthovel,andthereforeitstransportationinavesselisnotimprobable:analogousfactshavebeenobservedbyDr。
  RichardsoninNorthAmerica。
  Ofland-birdsIobtainedtwenty-sixkinds,allpeculiartothegroupandfoundnowhereelse,withtheexceptionofonelark-likefinchfromNorthAmericaDolichonyxoryzivorus,whichrangesonthatcontinentasfarnorthas54degs。,andgenerallyfrequentsmarshes。Theothertwenty-fivebirdsconsist,firstly,ofahawk,curiouslyintermediateinstructurebetweenabuzzardandtheAmericangroupofcarrion-feedingPolybori;andwiththeselatterbirdsitagreesmostcloselyineveryhabitandeventoneofvoice。Secondly,therearetwoowls,representingtheshort-earedandwhitebarn-owlsofEurope。Thirdly,awren,threetyrant-flycatcherstwoofthemspeciesofPyrocephalus,oneorbothofwhichwouldberankedbysomeornithologistsasonlyvarieties,andadove——allanalogousto,butdistinctfrom,Americanspecies。Fourthly,aswallow,whichthoughdifferingfromthePrognepurpureaofbothAmericas,onlyinbeingratherdullercolored,smaller,andslenderer,isconsideredbyMr。Gouldasspecificallydistinct。Fifthly,therearethreespeciesofmockingthrush——aformhighlycharacteristicofAmerica。Theremainingland-birdsformamostsingulargroupoffinches,relatedtoeachotherinthestructureoftheirbeaks,shorttails,formofbodyandplumage:therearethirteenspecies,whichMr。Gouldhasdividedintofoursubgroups。Allthesespeciesarepeculiartothisarchipelago;andsoisthewholegroup,withtheexceptionofonespeciesofthesub-groupCactornis,latelybroughtfromBowIsland,intheLowArchipelago。OfCactornis,thetwospeciesmaybeoftenseenclimbingabouttheflowersofthegreatcactus-trees;butalltheotherspeciesofthisgroupoffinches,mingledtogetherinflocks,feedonthedryandsterilegroundofthelowerdistricts。Themalesofall,orcertainlyofthegreaternumber,arejetblack;andthefemaleswithperhapsoneortwoexceptionsarebrown。ThemostcuriousfactistheperfectgradationinthesizeofthebeaksinthedifferentspeciesofGeospiza,fromoneaslargeasthatofahawfinchtothatofachaffinch,andifMr。Gouldisrightinincludinghissub-group,Certhidea,inthemaingroupeventothatofawarbler。ThelargestbeakinthegenusGeospizaisshowninFig。1,andthesmallestinFig。3;butinsteadoftherebeingonlyoneintermediatespecies,withabeakofthesizeshowninFig。2,therearenolessthansixspecieswithinsensiblygraduatedbeaks。Thebeakofthesub-groupCerthidea,isshowninFig。4。ThebeakofCactornisis[picture]
  1。Geospizamagnirostris。2。Geospizafortis。3。Geospizaparvula。4。
  Certhideaolivasea。
  somewhatlikethatofastarling,andthatofthefourthsubgroup,Camarhynchus,isslightlyparrot-shaped。Seeingthisgradationanddiversityofstructureinonesmall,intimatelyrelatedgroupofbirds,onemightreallyfancythatfromanoriginalpaucityofbirdsinthisarchipelago,onespecieshadbeentakenandmodifiedfordifferentends。Inalikemanneritmightbefanciedthatabirdoriginallyabuzzard,hadbeeninducedheretoundertaketheofficeofthecarrion-feedingPolyborioftheAmericancontinent。
  Ofwadersandwater-birdsIwasabletogetonlyelevenkinds,andoftheseonlythreeincludingarailconfinedtothedampsummitsoftheislandsarenewspecies。Consideringthewanderinghabitsofthegulls,Iwassurprisedtofindthatthespeciesinhabitingtheseislandsispeculiar,butalliedtoonefromthesouthernpartsofSouthAmerica。Thefargreaterpeculiarityoftheland-birds,namely,twenty-fiveoutoftwenty-six,beingnewspecies,oratleastnewraces,comparedwiththewadersandweb-footedbirds,isinaccordancewiththegreaterrangewhichtheselatterordershaveinallpartsoftheworld。Weshallhereafterseethislawofaquaticforms,whethermarineorfreshwater,beinglesspeculiaratanygivenpointoftheearth’ssurfacethantheterrestrialformsofthesameclasses,strikinglyillustratedintheshells,andinalesserdegreeintheinsectsofthisarchipelago。
  Twoofthewadersarerathersmallerthanthesamespeciesbroughtfromotherplaces:theswallowisalsosmaller,thoughitisdoubtfulwhetherornotitisdistinctfromitsanalogue。Thetwoowls,thetwotyrant-catchersPyrocephalusandthedove,arealsosmallerthantheanalogousbutdistinctspecies,towhichtheyaremostnearlyrelated;ontheotherhand,thegullisratherlarger。Thetwoowls,theswallow,allthreespeciesofmocking-thrush,thedoveinitsseparatecoloursthoughnotinitswholeplumage,theTotanus,andthegull,arelikewiseduskiercolouredthantheiranalogousspecies;andinthecaseofthemocking-thrushandTotanus,thananyotherspeciesofthetwogenera。Withtheexceptionofawrenwithafineyellowbreast,andofatyrant-flycatcherwithascarlettuftandbreast,noneofthebirdsarebrilliantlycoloured,asmighthavebeenexpectedinanequatorialdistrict。Henceitwouldappearprobable,thatthesamecauseswhichheremaketheimmigrantsofsomepeculiarspeciessmaller,makemostofthepeculiarGalapageianspeciesalsosmaller,aswellasverygenerallymoreduskycoloured。Alltheplantshaveawretched,weedyappearance,andIdidnotseeonebeautifulflower。Theinsects,again,aresmall-sizedanddull-coloured,and,asMr。Waterhouseinformsme,thereisnothingintheirgeneralappearancewhichwouldhaveledhimtoimaginethattheyhadcomefromundertheequator。[1]Thebirds,plants,andinsectshaveadesertcharacter,andarenotmorebrilliantlycolouredthanthosefromsouthernPatagonia;wemay,therefore,concludethattheusualgaudycolouringoftheinter-tropicalproductions,isnotrelatedeithertotheheatorlightofthosezones,buttosomeothercause,perhapstotheconditionsofexistencebeinggenerallyfavourabletolife。
  Wewillnowturntotheorderofreptiles,whichgivesthemoststrikingcharactertothezoologyoftheseislands。Thespeciesarenotnumerous,butthenumbersofindividualsofeachspeciesareextraordinarilygreat。
  ThereisonesmalllizardbelongingtoaSouthAmericangenus,andtwospeciesandprobablymoreoftheAmblyrhynchus——agenusconfinedtotheGalapagosIslands。Thereisonesnakewhichisnumerous;itisidentical,asIaminformedbyM。Bibron,withthePsammophisTemminckiifromChile。[2]Ofsea-turtleIbelievetherearemorethanonespecies,andoftortoisesthereare,asweshallpresentlyshow,twoorthreespeciesorraces。Oftoadsandfrogstherearenone:Iwassurprisedatthis,consideringhowwellsuitedforthemthetemperateanddampupperwoodsappearedtobe。ItrecalledtomymindtheremarkmadebyBorySt。
  Vincent,[3]namely,thatnoneofthisfamilyarefoundonanyofthevolcanicislandsinthegreatoceans。AsfarasIcanascertainfromvariousworks,thisseemstoholdgoodthroughoutthePacific,andeveninthelargeislandsoftheSandwicharchipelago。Mauritiusoffersanapparentexception,[m.kanbaapp.com]whereIsawtheRanaMascariensisinabundance:
  thisfrogissaidnowtoinhabittheSeychelles,Madagascar,andBourbon;
  butontheotherhand,DuBois,inhisvoyagein1669,statesthattherewerenoreptilesinBourbonexcepttortoises;andtheOfficierduRoiassertsthatbefore1768ithadbeenattempted,withoutsuccess,tointroducefrogsintoMauritius——Ipresumeforthepurposeofeating:henceitmaybewelldoubtedwhetherthisfrogisanaboriginaloftheseislands。Theabsenceofthefrogfamilyintheoceanicislandsisthemoreremarkable,whencontrastedwiththecaseoflizards,whichswarmonmostofthesmallestislands。Maythisdifferencenotbecaused,bythegreaterfacilitywithwhichtheeggsoflizards,protectedbycalcareousshellsmightbetransportedthroughsalt-water,thancouldtheslimyspawnoffrogs?
  IwillfirstdescribethehabitsofthetortoiseTestudonigra,formerlycalledIndica,whichhasbeensofrequentlyalludedto。Theseanimalsarefound,Ibelieve,onalltheislandsofthearchipelago;certainlyonthegreaternumber。Theyfrequentinpreferencethehighdampparts,buttheylikewiseliveinthelowerandariddistricts。Ihavealreadyshown,fromthenumberswhichhavebeencaughtinasingleday,howverynumeroustheymustbe。Somegrowtoanimmensesize:Mr。Lawson,anEnglishman,andvice-governorofthecolony,toldusthathehadseenseveralsolarge,thatitrequiredsixoreightmentoliftthemfromtheground;andthatsomehadaffordedasmuchastwohundredpoundsofmeat。Theoldmalesarethelargest,thefemalesrarelygrowingtosogreatasize:themalecanreadilybedistinguishedfromthefemalebythegreaterlengthofitstail。Thetortoiseswhichliveonthoseislandswherethereisnowater,orinthelowerandaridpartsoftheothers,feedchieflyonthesucculentcactus。Thosewhichfrequentthehigheranddampregions,eattheleavesofvarioustrees,akindofberrycalledguayavitawhichisacidandaustere,andlikewiseapalegreenfilamentouslichenUsneraplicata,thathangsfromtheboughsofthetrees。
  Thetortoiseisveryfondofwater,drinkinglargequantities,andwallowinginthemud。Thelargerislandsalonepossesssprings,andthesearealwayssituatedtowardsthecentralparts,andataconsiderableheight。Thetortoises,therefore,whichfrequentthelowerdistricts,whenthirsty,areobligedtotravelfromalongdistance。Hencebroadandwell-beatenpathsbranchoffineverydirectionfromthewellsdowntothesea-coast;andtheSpaniardsbyfollowingthemup,firstdiscoveredthewatering-places。WhenIlandedatChathamIsland,Icouldnotimaginewhatanimaltravelledsomethodicallyalongwell-chosentracks。Nearthespringsitwasacuriousspectacletobeholdmanyofthesehugecreatures,oneseteagerlytravellingonwardswithoutstretchednecks,andanothersetreturning,afterhavingdrunktheirfill。Whenthetortoisearrivesatthespring,quiteregardlessofanyspectator,heburieshisheadinthewaterabovehiseyes,andgreedilyswallowsgreatmouthfuls,attherateofaboutteninaminute。Theinhabitantssayeachanimalstaysthreeorfourdaysintheneighbourhoodofthewater,andthenreturnstothelowercountry;buttheydifferedrespectingthefrequencyofthesevisits。Theanimalprobablyregulatesthemaccordingtothenatureofthefoodonwhichithaslived。Itis,however,certain,thattortoisescansubsistevenontheseislandswherethereisnootherwaterthanwhatfallsduringafewrainydaysintheyear。
  Ibelieveitiswellascertained,thatthebladderofthefrogactsasareservoirforthemoisturenecessarytoitsexistence:suchseemstobethecasewiththetortoise。Forsometimeafteravisittothesprings,theirurinarybladdersaredistendedwithfluid,whichissaidgraduallytodecreaseinvolume,andtobecomelesspure。Theinhabitants,whenwalkinginthelowerdistrict,andovercomewiththirst,oftentakeadvantageofthiscircumstance,anddrinkthecontentsofthebladderiffull:inoneIsawkilled,thefluidwasquitelimpid,andhadonlyaveryslightlybittertaste。Theinhabitants,however,alwaysfirstdrinkthewaterinthepericardium,whichisdescribedasbeingbest。
  Thetortoises,whenpurposelymovingtowardsanypoint,travelbynightandday,andarriveattheirjourney’sendmuchsoonerthanwouldbeexpected。
  Theinhabitants,fromobservingmarkedindividuals,considerthattheytraveladistanceofabouteightmilesintwoorthreedays。Onelargetortoise,whichIwatched,walkedattherateofsixtyyardsintenminutes,thatis360yardsinthehour,orfourmilesaday,——allowingalittletimeforittoeatontheroad。Duringthebreedingseason,whenthemaleandfemalearetogether,themaleuttersahoarseroarorbellowing,which,itissaid,canbeheardatthedistanceofmorethanahundredyards。
  Thefemaleneveruseshervoice,andthemaleonlyatthesetimes;sothatwhenthepeoplehearthisnoise,theyknowthatthetwoaretogether。TheywereatthistimeOctoberlayingtheireggs。Thefemale,wherethesoilissandy,depositsthemtogether,andcoversthemupwithsand;butwherethegroundisrockyshedropsthemindiscriminatelyinanyhole:Mr。Bynoefoundsevenplacedinafissure。Theeggiswhiteandspherical;onewhichImeasuredwasseveninchesandthree-eighthsincircumference,andthereforelargerthanahen’segg。Theyoungtortoises,assoonastheyarehatched,fallapreyingreatnumberstothecarrion-feedingbuzzard。Theoldonesseemgenerallytodiefromaccidents,asfromfallingdownprecipices:
  atleast,severaloftheinhabitantstoldme,thattheyneverfoundonedeadwithoutsomeevidentcause。